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3. Anyone know more about Ephemerella septentrionalis?

I found about one sentence on these in Gonzo's book, and haven't seen them mentioned anywhere else in fly fishing literature. Nor is any of the scientific literature I have on them particularly interesting (just descriptions). Now that I've collected a few and see what unique-looking nymphs they've got, I'm really curious about them.

Jason,

I had a chance to talk with Penn State entomologist Greg Hoover yesterday. He said that he collects most of his septentrionalis nymphs by sweeping his net through mossy breaks on 1st and 2nd order streams. He also said that when he hears accounts of unusually large (three-tailed) "sulphurs," he suspects they may be this species. Apparently they are about a hook size larger than invaria on average. He confirmed that emergence timing is about the same as the early part of the invaria hatch. (I say "early part" because invaria can extend for quite a lengthy period on streams with heavy populations. The later, smaller component is even confused for dorothea on some streams that have no dorothea population.)

As you have noted, the easy field ID trait is the long spindly legs in both nymph and adult.

Thanks for getting that information. :)

Too bad I couldn't get a really intact dun. Still, given their rarity, I count myself lucky to have found this one.

Despite the very slight reference in fly-fishing literature, I'm not sure how rare they really are. They are rather widespread in the East, being found at least from Tennessee to Maine. The name implies they are "of the nothern regions," so I wouldn't be surprised if they spread well into Canada.

Even though Greg said he finds the highest populations in low order streams, the fact that your specimens came from much higher order streams indicates that at least token populations can be found in larger waters. Greg ran off quite a list of PA trout streams where he finds them, but their lesser significance compared to the overwhelming invaria populations is the reason I suggested they sometimes "support" those hatches. And even fairly bug-conscious fly fishers who encounter the hatch may not notice the differences between septentrionalis and invaria unless they collect specimens for comparison.

It may just be that they are more significant on smaller streams where matching the hatch is not very important, or that their habitat requirements are narrower than the very adaptable invaria. They are a very cool and interesting mayfly, and your photos of the nymph and dun are the only ones I've ever seen. And that does make your photos rare, indeed.

I'd suggest that the habitat requirements of septentrionalis may be narrower than those of invaria.

I just looked in Ernest Schwiebert's mega-volume Nymphs to see if he had anything about Ephemerella aurivillii (oddly, there's no mention of it), and I noticed he had some good info on septentrionalis, so I updated that hatch page with his information and some from this discussion. It's such a cool mayfly I figured I'd bump the topic.

Based on current taxonomy, Jason, my "sister sulphur" comment is inaccurate. Although some taxonomists included it in the "invaria group" in the past (Burks 1958), the only sense in which that comment applies today is that they may go to the same streamside dance together and look enough alike on the water that they might be mistaken by anglers. :)

Oh, and whether or not my book was the first mention of this species in the fly-fishing lit. might depend on what you mean by "mention." Charles Meck included this species in his hatch listing in Pennsylvania Trout Streams and Their Hatches (Backcountry Publications 1989), but I don't think he mentions anything about them in the main text. I believe that Charlie and I both learned to identify this species through discussions with Greg Hoover.

Okay, I took the "sister sulphur" part out. I still think it could be justified by the similarity in timing and appearance, with a caveat that it's not a taxonomical kinship, but I suppose it's better to avoid the confusion altogether.

See post-post edit above.

Jason,

This is a technical aside, but as I view this your actual comments and photos of the bug appear in a straight line down with only one word per line...Has anyone else had this problem or just me? All the posts seem ok just your original text and photos of the bug are all in one long line down.

Spence

Spence, I recall a similar bug on the front page for certain web browsers when I put this new update to the site online. Now I just have to remember how I fixed it.

I just thought you were showing us TroutNuts some examples of "Concrete Poetry"! ;)