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Artistic view of a Male Pteronarcys californica (Pteronarcyidae) (Giant Salmonfly) Stonefly Adult from the Gallatin River in Montana
Salmonflies
Pteronarcys californica

The giant Salmonflies of the Western mountains are legendary for their proclivity to elicit consistent dry-fly action and ferocious strikes.

Case view of a Pycnopsyche guttifera (Limnephilidae) (Great Autumn Brown Sedge) Caddisfly Larva from the Yakima River in Washington
It's only barely visible in one of my pictures, but I confirmed under the microscope that this one has a prosternal horn and the antennae are mid-way between the eyes and front of the head capsule.

I'm calling this one Pycnopsyche, but it's a bit perplexing. It seems to key definitively to at least Couplet 8 of the Key to Genera of Limnephilidae Larvae. That narrows it down to three genera, and the case seems wrong for the other two. The case looks right for Pycnopsyche, and it fits one of the key characteristics: "Abdominal sternum II without chloride epithelium and abdominal segment IX with only single seta on each side of dorsal sclerite." However, the characteristic "metanotal sa1 sclerites not fused, although often contiguous" does not seem to fit well. Those sclerites sure look fused to me, although I can make out a thin groove in the touching halves in the anterior half under the microscope. Perhaps this is a regional variation.

The only species of Pycnopsyche documented in Washington state is Pycnopsyche guttifera, and the colors and markings around the head of this specimen seem to match very well a specimen of that species from Massachusetts on Bugguide. So I am placing it in that species for now.

Whatever species this is, I photographed another specimen of seemingly the same species from the same spot a couple months later.
27" brown trout, my largest ever. It was the sub-dominant fish in its pool. After this, I hooked the bigger one, but I couldn't land it.
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This topic is about the Mayfly Genus Caudatella

This Western genus is unimportant, but Caudatella heterocaudata and Caudatella hystrix are said to sometimes produce fishable hatches.

Example specimens

Konchu
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Indiana

Posts: 498
Konchu on Oct 29, 2010October 29th, 2010, 1:36 pm EDT
The paper is in press now, so I can share an updated taxonomic synopsis of the "unimportant" genus Caudatella. Have fun.


Caudatella edmundsi (Allen, 1959)

Caudatella columbiella (McDunnough, 1935), comb. n.
= Ephemerella californica Allen and Edmunds, 1961, stat. n., syn. n.

Caudatella heterocaudata (McDunnough, 1929)
= Ephemerella circia Allen and Edmunds, 1961, stat. n., syn. n.

Caudatella hystrix (Traver, 1934)
= Ephemerella spinosa Mayo, 1952
= Ephemerella cascadia Allen and Edmunds, 1961

Caudatella jacobi (McDunnough, 1939)
= Ephemerella orestes Allen and Edmunds, 1961
Taxon
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Plano, TX

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Taxon on Oct 29, 2010October 29th, 2010, 3:18 pm EDT
Luke-

From your posting, it appears that what was previously Caudatella heterocaudata and its (3) subspecies are now Caudatella heterocaudata (with no subspecies), and (new species), Caudatella columbiella. Is that correct?
Best regards,
Roger Rohrbeck
www.FlyfishingEntomology.com
Konchu
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Indiana

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Konchu on Oct 30, 2010October 30th, 2010, 2:48 am EDT
columbiella was an old species that was synonymized with heterocaudata; i think it is a separate species

circia, californica and heterocaudata were all subspecies of heterocaudata

it turns out that californica is the same thing as columbiella

circia does not hold up as a nominal entity under most current philosophies. at the time it was described, the authors acknowledged that it probably interbred with other variants of heterocaudata, and the defining characters varied even in the population at the type locale. so, it should be considered the same species as heterocaudata
Taxon
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Plano, TX

Posts: 1311
Taxon on Oct 30, 2010October 30th, 2010, 10:42 am EDT
Thanks, Luke.
Best regards,
Roger Rohrbeck
www.FlyfishingEntomology.com

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