IDed these nymphs to genus using Merritt, Cummins and Berg (2008). The genus is monotypic. The only species in the genus is
Timpanoga hecuba which is divided into two subspecies,
Timpanoga hecuba hecuba and
Timpanoga hecuba pacifica. Personally, I think it should just be a single species. The nymphs of the two subspecies as described by Allen and Edmunds are separated by either having spines on the posterior edges of the abdominal tergites (
Timpanoga hecuba pacifica) or not (
Timpanoga hecuba hecuba). Collections I've gotten from Mill Creek (tributary of the Russian River) have the spines on about half the specimens while the others are entirely without. So far as I know only
Timpanoga hecuba pacifica has had the adults described.
The paper by Allen and Edmunds describing
Timpanoga hecuba and its division into the two subspecies is "A Revision of the Genus Ephemerella (Ephemeroptera: Ephemerellidae). I. The Subgenus Timpanoga" and can be found here:
http://www.ephemeroptera-galactica.com/pubs/pub_a/puballenr1959p51.pdf
The nymphs are most common from mid-summer through late fall and are found in slack water at the edges of pools or glides. They often have a coating of silt on them.