The giant Salmonflies of the Western mountains are legendary for their proclivity to elicit consistent dry-fly action and ferocious strikes.
"The clay substrate inhabited by Tortopus is always firm, nearly always vertical, and usually exposed to swift water."
McCafferty (1975) collected nymphs in hard clay banks approximitely three feet below the water surface in the Wabash River in Indiana. He found the clay banks to be honeycombed by Pentagenia nymphs in the same manner that the banks of the Savannah River are honeycombed by Tortopus.