This Skwala nymph still has a couple months left to go before hatching, but it's still a good representative of its species, which was extremely abundant in my sample for a stonefly of this size. It's obvious why the Yakima is known for its Skwala hatch.
Many of America's traditional flies, like the March Brown and the Light and Dark Cahills, originated in the Catskills to imitate the mayflies of this genus. Caucci and Nastasi wrote of them in Hatches II:
"There is a matrimony between Stenonema flies and Catskill rivers that is as synonymous as ham and eggs."
By far the most important species is Stenonema vicarium, variants of which are known to anglers as the March Brown and Gray Fox. Of the others in the "Cahill" group, Stenonema ithaca is the next most important, along with Stenonema modestum and Stenonema pulchellum, which also produce localized fishable hatches. These lesser species together with a few species of Stenacron and the Stenonema femoratum represent the sporadic Light Cahill hatches on evenings in late Spring that often continue throughout the Summer.
Entoman on May 21, 2012May 21st, 2012, 8:03 am EDT
Thanks Jeff, for the excellent photo and determination. Readers note that besides the three spots that are clearly seen, spots are also just visible on seg's 3 and 8. The dark transverse dashes are a little harder to make out, but they look to be present as well. The crowded crossveins below the bulla are also indicative of femoratum. This is the last Stenonema species left in what is now the least important genus (to anglers) of the related group. See the genus topic article as well as the family article in the hatch encyclopedia for a brief explanation.
"It's not that I find fishing so important, it's just that I find all other endeavors of Man equally unimportant... And not nearly as much fun!" Robert Traver, Anatomy of a Fisherman