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Lateral view of a Female Hexagenia limbata (Ephemeridae) (Hex) Mayfly Dun from the Namekagon River in Wisconsin
Hex Mayflies
Hexagenia limbata

The famous nocturnal Hex hatch of the Midwest (and a few other lucky locations) stirs to the surface mythically large brown trout that only touch streamers for the rest of the year.

Dorsal view of a Grammotaulius betteni (Limnephilidae) (Northern Caddisfly) Caddisfly Larva from the Yakima River in Washington
This is a striking caddis larva with an interesting color pattern on the head. Here are some characteristics I was able to see under the microscope, but could not easily expose for a picture:
- The prosternal horn is present.
- The mandible is clearly toothed, not formed into a uniform scraper blade.
- The seems to be only 2 major setae on the ventral edge of the hind femur.
- Chloride epithelia seem to be absent from the dorsal side of any abdominal segments.
Based on these characteristics and the ones more easily visible from the pictures, this seems to be Grammotaulius. The key's description of the case is spot-on: "Case cylindrical, made of longitudinally arranged sedge or similar leaves," as is the description of the markings on the head, "Dorsum of head light brownish yellow with numerous discrete, small, dark spots." The spot pattern on the head is a very good match to figure 19.312 of Merritt R.W., Cummins, K.W., and Berg, M.B. (2019). The species ID is based on Grammotaulius betteni being the only species of this genus known in Washington state.
27" brown trout, my largest ever. It was the sub-dominant fish in its pool. After this, I hooked the bigger one, but I couldn't land it.
Troutnut is a project started in 2003 by salmonid ecologist Jason "Troutnut" Neuswanger to help anglers and fly tyers unabashedly embrace the entomological side of the sport. Learn more about Troutnut or support the project for an enhanced experience here.

This topic is about the Mayfly Genus Epeorus

There is remarkable variety of form and color within this prolific genus of fast-water mayflies. Different species are found across the country, and several cause good hatches. Fly anglers are likely to encounter the lesser species on occasion, too.

The best Epeorus hatch in the East is Epeorus pleuralis, the famous Quill Gordon, the first abundant large mayfly hatch of the year. Epeorus vitreus comes a little later and is important in both the East and Midwest.

In the West, Epeorus longimanus dominates in fast, high-altitude streams, while Epeorus albertae inhabits slower and lower waters.

Example specimens

GONZO
Site Editor
"Bear Swamp," PA

Posts: 1681
GONZO on Oct 4, 2006October 4th, 2006, 7:27 am EDT
I believe rubidus = vitreus
Troutnut
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Administrator
Bellevue, WA

Posts: 2758
Troutnut on Oct 4, 2006October 4th, 2006, 7:51 am EDT
Thanks. Good catch. That must be a recent change, or at least recently added to the mayfly species list at Purdue.

I have one small thing on my programming to-do list before I can make the change, but I'll do it soon.
Jason Neuswanger, Ph.D.
Troutnut and salmonid ecologist
Taxon
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Site Editor
Plano, TX

Posts: 1311
Taxon on Oct 4, 2006October 4th, 2006, 8:08 am EDT
... That must be a recent change, or at least recently added to the mayfly species list at Purdue. ...


Jason-

Right you are. These are the most recent updates.
Best regards,
Roger Rohrbeck
www.FlyfishingEntomology.com
Troutnut
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Administrator
Bellevue, WA

Posts: 2758
Troutnut on Oct 4, 2006October 4th, 2006, 8:12 am EDT
Thanks for compiling that list! It will be a big time-saver when I'm ready to update my taxa.
Jason Neuswanger, Ph.D.
Troutnut and salmonid ecologist

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