This Skwala nymph still has a couple months left to go before hatching, but it's still a good representative of its species, which was extremely abundant in my sample for a stonefly of this size. It's obvious why the Yakima is known for its Skwala hatch.
This Western genus is closely related to Epeorus and its biology is similar. The only species recognized as important to fly anglers is Ironodes nitidus.
Where & when
In 10 records from GBIF, adults of this genus have been collected during June (60%), May (20%), March (10%), and July (10%).
In 5 records from GBIF, this genus has been collected at elevations ranging from 450 to 5200 ft, with an average (median) of 3600 ft.
While relatively rare in most of western Montana, it is common in several small streams along the east side of Flathead Lake and streams on the south side of the Flathead Indian Reservation.