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Baetis
Tiny Baetis mayflies are perhaps the most commonly encountered and imitated by anglers on all American trout streams due to their great abundance, widespread distribution, and trout-friendly emergence habits.
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It's only barely visible in one of my pictures, but I confirmed under the microscope that this one has a prosternal horn and the antennae are mid-way between the eyes and front of the head capsule.
I'm calling this one Pycnopsyche, but it's a bit perplexing. It seems to key definitively to at least Couplet 8 of the Key to Genera of Limnephilidae Larvae. That narrows it down to three genera, and the case seems wrong for the other two. The case looks right for Pycnopsyche, and it fits one of the key characteristics: "Abdominal sternum II without chloride epithelium and abdominal segment IX with only single seta on each side of dorsal sclerite." However, the characteristic "metanotal sa1 sclerites not fused, although often contiguous" does not seem to fit well. Those sclerites sure look fused to me, although I can make out a thin groove in the touching halves in the anterior half under the microscope. Perhaps this is a regional variation.
The only species of Pycnopsyche documented in Washington state is Pycnopsyche guttifera, and the colors and markings around the head of this specimen seem to match very well a specimen of that species from Massachusetts on Bugguide. So I am placing it in that species for now.
Whatever species this is, I photographed another specimen of seemingly the same species from the same spot a couple months later.
I'm calling this one Pycnopsyche, but it's a bit perplexing. It seems to key definitively to at least Couplet 8 of the Key to Genera of Limnephilidae Larvae. That narrows it down to three genera, and the case seems wrong for the other two. The case looks right for Pycnopsyche, and it fits one of the key characteristics: "Abdominal sternum II without chloride epithelium and abdominal segment IX with only single seta on each side of dorsal sclerite." However, the characteristic "metanotal sa1 sclerites not fused, although often contiguous" does not seem to fit well. Those sclerites sure look fused to me, although I can make out a thin groove in the touching halves in the anterior half under the microscope. Perhaps this is a regional variation.
The only species of Pycnopsyche documented in Washington state is Pycnopsyche guttifera, and the colors and markings around the head of this specimen seem to match very well a specimen of that species from Massachusetts on Bugguide. So I am placing it in that species for now.
Whatever species this is, I photographed another specimen of seemingly the same species from the same spot a couple months later.
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Quick evening on the upper Yakima
Quick evening on the upper Yakima
By Troutnut on September 16th, 2020
With smoke from the west coast wildfires beginning to clear just a little, I took a few hours to fish one of the nearest access points on the Yakima River, where the fish are small and the water uncrowded. There were some stoneflies and caddisflies in the air--I collected one of each, representing the most abundant species--but there weren't enough bugs on the water to get the fish rising, except for the abundant 3-inch-long Chinook Salmon parr. I continued playing around with my new Euro nymphing rig and landed several rainbows up to 12", including my first double on that rod.
Photos by Troutnut from the Yakima River in Washington
Closeup insects by Troutnut from the Yakima River in Washington
I first just assumed this was Dicosmoecus based on anglers' conventional wisdom since it's a large orange "October caddis," but Creno set me straight. I should have keyed it out. After another look under the microscope, it lacks an anepisternal wart on the mesopleuron, which rules out Dicosmoecus. The midtibiae have 2 apical spurs and 1 pre-apical spur, and from there the color pattern of the wing points to Onocosmoecus. The location then narrows the species to unicolor.
The family ID on this one was a little bit tricky. Just going by the size, shape, and color, it looks like Chloroperlidae. However, the second anal vein of the forewing is does not appear to be forked, and the apical maxillary palpal segment is close to the length of the penultimate segment, both of which rule out that family. The position of the cubitoanal crossvein relative to the anal cell in the forewing -- touching it in this case -- indicates Perlidae (and it really doesn't have the "look" of Perlidae at all), but other characteristics, such as the metathorastic sternacostal sutures and lack of gill remnants, point to Perlodidae. That's the right answer. Moving on to Perlodidae, the key characteristics in Merritt & Cummins lead straightforwarly to Isoperla, and the species key in Jewett 1959 (The Stoneflies of the Pacific Northwest) leads to Isoperla fusca.
There is one caveat: That source does suggest a May-July emergence, whereas this one was collected in mid-September.
There is one caveat: That source does suggest a May-July emergence, whereas this one was collected in mid-September.
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