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Lateral view of a Male Baetis (Baetidae) (Blue-Winged Olive) Mayfly Dun from Mystery Creek #43 in New York
Blue-winged Olives
Baetis

Tiny Baetis mayflies are perhaps the most commonly encountered and imitated by anglers on all American trout streams due to their great abundance, widespread distribution, and trout-friendly emergence habits.

Dorsal view of a Sweltsa (Chloroperlidae) (Sallfly) Stonefly Nymph from the Yakima River in Washington
This species was fairly abundant in a February sample of the upper Yakima.
27" brown trout, my largest ever. It was the sub-dominant fish in its pool. After this, I hooked the bigger one, but I couldn't land it.
Troutnut is a project started in 2003 by salmonid ecologist Jason "Troutnut" Neuswanger to help anglers and fly tyers unabashedly embrace the entomological side of the sport. Learn more about Troutnut or support the project for an enhanced experience here.

Sulphur Duns

Like most common names,"Sulphur Dun" can refer to more than one taxon. They're previewed below, along with 5 specimens. For more detail click through to the scientific names.

Mayfly Species Ephemerella dorothea dorothea

These are often called Sulphur Duns.
Ephemerella dorothea consists of two subspecies, which both produce excellent action. Ephemerella dorothea dorothea is a small species of Sulphur in the East, and Ephemerella dorothea infrequens (formerly Ephemerella infrequens) is one of the two main Pale Morning Dun hatches of the West. The remainder of this page focuses on the dorothea dorothea subspecies, and Ephemerella dorothea infrequens is discussed separately on its own page.

This is one of the most challenging mayfly hatches on Eastern waters. On many streams, it follows or overlaps hatches of the larger, lingering Ephemerella invaria.
Dorsal view of a Ephemerella dorothea dorothea (Ephemerellidae) (Pale Evening Dun) Mayfly Nymph from Paradise Creek in Pennsylvania
I keyed this nymph carefully under a microscope to check that it's Ephemerella dorothea.

Mayfly Species Ephemerella invaria

These are often called Sulphur Duns.
This species, the primary "Sulphur" hatch, stirs many feelings in the angler. There is nostalgia for days when everything clicked and large, selective trout were brought to hand. There is the bewildering memory of towering clouds of spinners which promise great fishing and then vanish back into the aspens as night falls. There is frustration from the maddening selectivity with which trout approach the emerging duns--a vexing challenge that, for some of us, is the source of our excitement when Sulphur time rolls around.

Ephemerella invaria is one of the two species frequently known as Sulphurs (the other is Ephemerella dorothea). There used to be a third, Ephemerella rotunda, but entomologists recently discovered that invaria and rotunda are a single species with an incredible range of individual variation. This variation and the similarity to the also variable dorothea make telling them apart exceptionally tricky.

As the combination of two already prolific species, this has become the most abundant of all mayfly species in Eastern and Midwestern trout streams.
Lateral view of a Male Ephemerella invaria (Ephemerellidae) (Sulphur) Mayfly Dun from Penn's Creek in Pennsylvania
Lateral view of a Male Ephemerella invaria (Ephemerellidae) (Sulphur) Mayfly Spinner from the Teal River in Wisconsin
Dorsal view of a Ephemerella invaria (Ephemerellidae) (Sulphur) Mayfly Nymph from the Beaverkill River in New York
This small Ephemerella invaria nymph was at least a month away from emergence.

Sulphur Duns

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