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Lateral view of a Female Hexagenia limbata (Ephemeridae) (Hex) Mayfly Dun from the Namekagon River in Wisconsin
Hex Mayflies
Hexagenia limbata

The famous nocturnal Hex hatch of the Midwest (and a few other lucky locations) stirs to the surface mythically large brown trout that only touch streamers for the rest of the year.

Dorsal view of a Grammotaulius betteni (Limnephilidae) (Northern Caddisfly) Caddisfly Larva from the Yakima River in Washington
This is a striking caddis larva with an interesting color pattern on the head. Here are some characteristics I was able to see under the microscope, but could not easily expose for a picture:
- The prosternal horn is present.
- The mandible is clearly toothed, not formed into a uniform scraper blade.
- The seems to be only 2 major setae on the ventral edge of the hind femur.
- Chloride epithelia seem to be absent from the dorsal side of any abdominal segments.
Based on these characteristics and the ones more easily visible from the pictures, this seems to be Grammotaulius. The key's description of the case is spot-on: "Case cylindrical, made of longitudinally arranged sedge or similar leaves," as is the description of the markings on the head, "Dorsum of head light brownish yellow with numerous discrete, small, dark spots." The spot pattern on the head is a very good match to figure 19.312 of Merritt R.W., Cummins, K.W., and Berg, M.B. (2019). The species ID is based on Grammotaulius betteni being the only species of this genus known in Washington state.
27" brown trout, my largest ever. It was the sub-dominant fish in its pool. After this, I hooked the bigger one, but I couldn't land it.
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This topic is about the Caddisfly Genus Chimarra

Chimarra aterrima is the most important species.

Example specimens

Btopbuckeye
Btopbuckeye's profile picture
Hayesville no

Posts: 1
Btopbuckeye on Mar 4, 2015March 4th, 2015, 3:57 pm EST
I had a large swarm of black sedges in a size 16 ovipositing today.the females hadvblack wings and body with a green egg sack was just wondering if that is this genus or something totally different
Entoman
Entoman's profile picture
Northern CA & ID

Posts: 2604
Entoman on Mar 5, 2015March 5th, 2015, 7:29 am EST
Welcome to the forum. They are most likely Brachycentrus (Grannoms).
"It's not that I find fishing so important, it's just that I find all other endeavors of Man equally unimportant... And not nearly as much fun!" Robert Traver, Anatomy of a Fisherman
Gutcutter
Gutcutter's profile picture
Pennsylvania

Posts: 470
Gutcutter on Mar 5, 2015March 5th, 2015, 7:50 am EST
Are you sure they were caddis and not stoneflies?
The larger "little black stones" (around a size 16) start to flutter around this time of year up here.
The egg sacks I see are yellowish on the stones.
I'm not sure of the timing of Grannoms or stones in N.C.
All men who fish may in turn be divided into two parts: those who fish for trout and those who don't. Trout fishermen are a race apart: they are a dedicated crew- indolent, improvident, and quietly mad.

-Robert Traver, Trout Madness
Crepuscular
Crepuscular's profile picture
Boiling Springs, PA

Posts: 920
Crepuscular on Mar 5, 2015March 5th, 2015, 9:10 am EST
Welcome to the forum. They are most likely Brachycentrus (Grannoms).


I was going to say the same Kurt but the description of the Black Wings had me wondering, really Btopbuckeye, It's virtually impossible to say with any kind of certainty what they may have been. But as Kurt pointed out they may have been Brachycentrus sp., the green eggs would be consistent with that. And I imagine you are a head of us here in PA with timing of hatches and we usually see Brachycentrus (Grannoms) in mid to late April here in southern PA. I've heard those Rainbows in the Nantahalla are beautiful fish! I'd like to see one up close and personal some day.
Martinlf
Martinlf's profile picture
Moderator
Palmyra PA

Posts: 3047
Martinlf on Mar 5, 2015March 5th, 2015, 11:56 am EST
Given that the egg sacks were green, it would seem they were caddis of some sort. At least one hatch chart for the Smokies lists "Grannoms" this early:

http://www.perfectflystore.com/hcsmokymtns.html

And they provide specific information about the bug, which they call a "little black caddis":

http://www.perfectflystore.com/littleblcadisinfo.html

They identify the Grannom that hatches early in the Smokies as Brachycentrus occidentalis.

In regard to our Grannoms here in PA, one of Troutnut's bug experts, Lloyd Gonzales observes: "The collecting record shows two Brachycentrus spp. for Huntingdon County [PA]--B. numerosus and solomoni"; and he identifies numerosus as the species most commonly found on Penns Creek.

Eric, might the possible species difference at least partially explain the difference in hatch time, along with geographical differences?

Brian, The Nantahala is certainly a beautiful river; along with the Ocoee it was a favorite when I was a whitewater guide in North Carolina. We primarily frequented the lower reaches, taking out just after the first falls above the Nantahala Outdoor Center. I once saw a kid walking along one of the feeder streams to the Nantahala carrying a stringer of big rainbows. Obviously not a proponent of catch and release.
"He spread them a yard and a half. 'And every one that got away is this big.'"

--Fred Chappell
Oldredbarn
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Novi, MI

Posts: 2600
Oldredbarn on Mar 5, 2015March 5th, 2015, 1:50 pm EST
Welcome to the forum. They are most likely Brachycentrus (Grannoms).


Kurt...Every darn hatch chart in this state still has the "Little Black Caddis" listed as Chimarra! And listed as an 16/18 hook!? Even after Carl Richards, no less, told them about, B lateralis, B numerosus, and B americanus...

I have made loud, Molson infused, bets to some folk here that they probably haven't ever seen a Chimarra.

Spence
"Even when my best efforts fail it's a satisfying challenge, and that, after all, is the essence of fly fishing." -Chauncy Lively

"Envy not the man who lives beside the river, but the man the river flows through." Joseph T Heywood
Crepuscular
Crepuscular's profile picture
Boiling Springs, PA

Posts: 920
Crepuscular on Mar 6, 2015March 6th, 2015, 3:41 am EST
Eric, might the possible species difference at least partially explain the difference in hatch time, along with geographical differences?


yes

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