Artistic view of a Male Pteronarcys californica (Pteronarcyidae) (Giant Salmonfly) Stonefly Adult from the Gallatin River in Montana

Salmonflies
Pteronarcys californica

The giant Salmonflies of the Western mountains are legendary for their proclivity to elicit consistent dry-fly action and ferocious strikes.

View from atop the permafrost slump

From the Selawik River in Alaska

Stream photos

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Tricorythodes (Trico) Mayfly Nymph Tricorythodes (Trico) Mayfly Nymph

Dorsal view of a Tricorythodes (Leptohyphidae) (Trico) Mayfly Nymph from Mystery Creek #308 in Washington

9 Recently Added Photos

27" brown trout, my largest ever. It was the sub-dominant fish in its pool. After this, I hooked the bigger one, but I couldn't land it.

Troutnut is a project started in 2003 by salmonid ecologist Jason "Troutnut" Neuswanger to help anglers and fly tyers unabashedly embrace the entomological side of the sport. Learn more about Troutnut or support the project for an enhanced experience here.

Agapetus are EVERYWHERE!!!!

This topic is about the Caddisfly Family Glossosomatidae

This family is one of the primitive caddisflies of the Rhyacophiloidea superfamily. However, they are not free-living like their better known cousins the Rhyacophilidae (Green Rockworm). Instead they build rounded "turtle" shaped cases that do not surround the larvae but are rather attached to the rock surface at their margins. Underneath is a sling made of secretions upon which the larvae ride, hence the common name Saddle-case Makers. There are four genera of possible interest but only one is generally recognized as important to anglers. See Glossosoma (Little Brown Short-horned Caddis) for details. The other three are so tiny that they are also called Pseudo-microcaddis.

Protoptila (Tiny Spotted Short-horned Caddis) is rarely important in trout streams and is generally found in warmer, larger water than the other two genera.

Agapetus (Tiny black Short-horned Caddis) is quite common in many northern streams.

Matrioptila is an extremely tiny southern genus.

Example specimens

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Male Glossosoma (Little Brown Short-horned Sedge) Caddisfly Adult

Lateral view of a Male Glossosoma (Glossosomatidae) (Little Brown Short-horned Sedge) Caddisfly Adult from the Cedar River in Washington

Key characteristics verified under the microscope: -The head has 3 ocelli. -Maxillary palp has 5 segments, with the second segment rounded. In some of the pictures it looks like 6 with the 3rd segment rounded, but the basal "segment" in that case isn't part of the palp. -Foretibiae lack preapical spurs. -Pronotum with mesal setose warts widely spaced. All of this very clearly indicates Glossosomatidae. Beyond there: -The tibial spur counts (fore / middle / hind tibiae) are 2, 4, and 4. -From the pictures, the hind wing discoidal cell is closed, not open. Subfamily Glossosomatinae. -The upper and lower parts of the mesepisternum are separated by a constriction, not a transverse suture. Microscope picture included. This leads to a very confident genus ID of Glossosoma. I don't have the information to take it to species.

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Glossosoma (Little Brown Short-horned Sedge) Caddisfly Larva

Dorsal view of a Glossosoma (Glossosomatidae) (Little Brown Short-horned Sedge) Caddisfly Larva from the Yakima River in Washington

I caught this tiny larva without a case, but it seems to key pretty clearly to to Glossosomatidae. From there, the lack of sclerites on the mesonotum points to either Glossosoma or Anagapetus. Although it's difficult to see in a 2D image from the microscope, it's pretty clear in the live 3D view that the pronotum is only excised about 1/3 of its length to accommodate the forecoxa, not 2/3, which points to Glossosoma at Couplet 5 of the Key to Genera of Glossosomatidae Larvae.

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Glossosoma (Little Brown Short-horned Sedge) Caddisfly Adult

Glossosoma (Glossosomatidae) (Little Brown Short-horned Sedge) Caddisfly Adult from the Big Thompson River in Montana

Litobrancha
Knoxville TN

Posts: 51

Litobrancha on Apr 12, 2007April 12th, 2007, 3:41 pm EDT

Jason

just wanted to spread the word about agapetus. many trout streams have healthy populations of agapetus and there is no reason that some of these species are important to early season emerger/dry fly fishing. small (#18-22) black caddis dry or emerger patterns will mimic them nicely, as well as Dolophilodes Wormaldia and Chimarra.

my colleagues are describing 12 new species of agapetus, mostly from the southeastern united states. i would encourage troutnuts to attempt to collect and rear agapetus pupae. it is pretty easy to do, find pupating cases and remove them from the rocks using forceps and into a small jar of water. If you use a jar with a small amount of water (just a little bit more than required to cover the pupae, removing the small stones around the puparium), then they will pupate in a refrigerator (preferably 60 C or so). Leave the lid loose to allow oxygen to equilibrate with the pupae. This also works for Rhyacophila, which build a similar puparium. We are describing new species of both Agapetus and Rhyacophila and it would be great to have specimens from Troutnuts!!! if interested in doing this, and it is time, email me litobrancha@yahoo.com You can send them to me in alcohol, who knows what else is out there!!!!

GONZO
Site Editor
"Bear Swamp," PA

Posts: 1681

GONZO on Apr 12, 2007April 12th, 2007, 6:04 pm EDT

Great, Lito, another genus to add to my tying list--just what I needed! :) Just kidding pal, it's great to hear from you again! And I'll be on the lookout for Agapetus! (Actually, if Chimarra/Dolophilodes imitations work, I'm all set.) ;)

Thanks,
Lloyd