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Lateral view of a Male Baetis (Baetidae) (Blue-Winged Olive) Mayfly Dun from Mystery Creek #43 in New York
Blue-winged Olives
Baetis

Tiny Baetis mayflies are perhaps the most commonly encountered and imitated by anglers on all American trout streams due to their great abundance, widespread distribution, and trout-friendly emergence habits.

Dorsal view of a Neoleptophlebia (Leptophlebiidae) Mayfly Nymph from the Yakima River in Washington
Some characteristics from the microscope images for the tentative species id: The postero-lateral projections are found only on segment 9, not segment 8. Based on the key in Jacobus et al. (2014), it appears to key to Neoleptophlebia adoptiva or Neoleptophlebia heteronea, same as this specimen with pretty different abdominal markings. However, distinguishing between those calls for comparing the lengths of the second and third segment of the labial palp, and this one (like the other one) only seems to have two segments. So I'm stuck on them both. It's likely that the fact that they're immature nymphs stymies identification in some important way.
27" brown trout, my largest ever. It was the sub-dominant fish in its pool. After this, I hooked the bigger one, but I couldn't land it.
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This topic is about the Caddisfly Genus Dicosmoecus

This is a superhatch in the West. Gary LaFontaine had a self-proclaimed obsession with Dicosmoecus, and he devotes several pages of Caddisflies to stressing the importance of its larvae, pupae, and adults. He wrote:

The question for fly fishermen seeking big trout is: "Which insects provide the best opportunity for catching such fish?" My list would be: Giant Orange Sedge (Dicosmoecus sp.), Salmon Fly (Pteronarcys californica, a stonefly), and the Michigan Mayfly (Hexagenia limbata). Dicosmoecus is the most important -- and the contest is not even close.

His reasons for this judgement are five traits of Dicosmoecus:

  • Very large size: Adults 30mm long, and thick.

  • Activity concentrated within 2-3 weeks.

  • Emerges in the low, clear water of fall.

  • Active during afternoon and evening.

  • Abundant in rivers with sea-run trout.


Personally I have only caught a good Dicosmoecus hatch one time, and on a secret desert spring creek. It reminded me of fishing Isonychia mayfly hatches back east, because there weren't enough bugs around to have trout rising consistently, but the presence of a particularly good food source in low numbers for a long time had them on the lookout for imitations. I had big trout cartwheeling out of the water to smash actively-fished imitations.

Example specimens

Troutnut
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Administrator
Bellevue, WA

Posts: 2758
Troutnut on Aug 22, 2006August 22nd, 2006, 3:42 pm EDT
There's a good topic with people sharing their experiences with Dicosmoecus over at the Washington Fly Fishing Forum.
Jason Neuswanger, Ph.D.
Troutnut and salmonid ecologist
Taxon
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Site Editor
Plano, TX

Posts: 1311
Taxon on Aug 22, 2006August 22nd, 2006, 7:40 pm EDT
Jason-

In case you're not familiar with the cast of characters on WFF, that particular thread drew several pretty savvy members. Preston is Preston Singletary a columnist with Flyfishing & Tying Journal. Smamla (as in S. malma) is Curt Kramer, a retired WDFW fisheries biologist and renowned Bull Trout enthusiast.
Best regards,
Roger Rohrbeck
www.FlyfishingEntomology.com

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