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Artistic view of a Male Pteronarcys californica (Pteronarcyidae) (Giant Salmonfly) Stonefly Adult from the Gallatin River in Montana
Salmonflies
Pteronarcys californica

The giant Salmonflies of the Western mountains are legendary for their proclivity to elicit consistent dry-fly action and ferocious strikes.

Case view of a Pycnopsyche guttifera (Limnephilidae) (Great Autumn Brown Sedge) Caddisfly Larva from the Yakima River in Washington
It's only barely visible in one of my pictures, but I confirmed under the microscope that this one has a prosternal horn and the antennae are mid-way between the eyes and front of the head capsule.

I'm calling this one Pycnopsyche, but it's a bit perplexing. It seems to key definitively to at least Couplet 8 of the Key to Genera of Limnephilidae Larvae. That narrows it down to three genera, and the case seems wrong for the other two. The case looks right for Pycnopsyche, and it fits one of the key characteristics: "Abdominal sternum II without chloride epithelium and abdominal segment IX with only single seta on each side of dorsal sclerite." However, the characteristic "metanotal sa1 sclerites not fused, although often contiguous" does not seem to fit well. Those sclerites sure look fused to me, although I can make out a thin groove in the touching halves in the anterior half under the microscope. Perhaps this is a regional variation.

The only species of Pycnopsyche documented in Washington state is Pycnopsyche guttifera, and the colors and markings around the head of this specimen seem to match very well a specimen of that species from Massachusetts on Bugguide. So I am placing it in that species for now.

Whatever species this is, I photographed another specimen of seemingly the same species from the same spot a couple months later.
27" brown trout, my largest ever. It was the sub-dominant fish in its pool. After this, I hooked the bigger one, but I couldn't land it.
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Female Dicosmoecus atripes (October Caddis) Caddisfly Adult Pictures

This specimen is 26 mm.
- Entoman

This caddisfly was collected from the St. Joe River in Idaho on September 27th, 2013 and added to Troutnut.com by Entoman on September 29th, 2013.

Discussions of this Adult

Gilvipes or atripes?
7 replies
Posted by Entoman on Sep 29, 2013
Last reply on Dec 19, 2016 by Clark2016
These two species are very difficult to tell apart. Because of the fuscated wing veining and black antennae, I'm leaning towards gilvipes, but it is my understanding that these characters are not very dependable. Regardless, both are the October caddis of western lore and can provide awesome dry fly fishing if the angler is lucky enough to hit it right.

There is some debate over which flies work better, dries or nymphs. I come down on the side of dry flies. I have rarely found the pupa effective, and even when they are it's often a darker color than the conventional orange that works best. For years I struggled with this until learning that the pupa (more properly called pharates) usually crawl out on shore to ecclode in the darkness of night. Since they also start from their rock cabins in very shallow water to begin with - not much nymphing opportunity there... It is highly doubtful that trout have the opportunity to feed much on the pharates, let alone enough to become selective. It is my opinion that when the pupal imitations work well, it is because they are either taken for drowned adults or perhaps it's just an attractor response to what looks like a big chunk of protein. Big fish often lose their caution this time of year as fishing pressure abates and storing calories for the Winter becomes a prerequisite.

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Female Dicosmoecus atripes (October Caddis) Caddisfly Adult Pictures

Collection details
Location: St. Joe River, Idaho
Date: September 27th, 2013
Added to site: September 29th, 2013
Author: Entoman
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