Tiny Baetis mayflies are perhaps the most commonly encountered and imitated by anglers on all American trout streams due to their great abundance, widespread distribution, and trout-friendly emergence habits.
This wild-looking little thing completely puzzled me. At first I was thinking beetle or month larva, until I got a look at the pictures on the computer screen. I made a couple of incorrect guesses before entomologist Greg Courtney pointed me in the right direction with Psychodidae. He suggested a possible genus of Thornburghiella, but could not rule out some other members of the tribe Pericomini.
Ephemerella dorothea infrequens (formerly Ephemerella infrequens), together with its often smaller and later hatching sibling Ephemerella excrucians, make up the most important Western hatches. They go by several common names but are best known as Pale Morning Duns (PMD's). They are rivaled only by the many baetid species that typically bookend them. In terms of availability, consistency and abundance (not to mention their convenient timing and preference for beautiful weather), they have no rival. They can run in size from a large 14 to a small 16 and various shades of illusive yellowish creams, sulfurs, and even yellowish greens, depending on the system they inhabit.
This taxon used to be considered the separate species Ephemerella infrequens, but entomologists now regard it as a subspecies together with the small eastern Pale Evening Dun hatch Ephemerella dorothea dorothea. There is another related listing of significance common in California and the Southwest that has undergone revision. The large (often exceeding 10 mm) Ephemerella mollitia is now considered synonymous with d. infrequens.
While it is not the normal policy of TroutNut to list subspecies as separate taxa, d. infrequens and d. dorothea are so important and distinct from each other in terms of geography, appearence, and angling tradition that they warrant an exception.