Salmonflies
Pteronarcys californica
The giant Salmonflies of the Western mountains are legendary for their proclivity to elicit consistent dry-fly action and ferocious strikes.
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This dun emerged from a mature nymph on my desk. Unfortunately its wings didn't perfectly dry out.
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Identification: Key to Orders of Adult Aquatic Insects, Couplet 5
Identification: Key to Orders of Adult Aquatic Insects, Couplet 5
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Adapted from Merritt R.W., Cummins, K.W., and Berg, M.B. (2019)
| Option 1 | Option 2 |
|---|---|
| Two or three tails | No tails (conspicuous filaments at the end of the abdomen) |
| Mouthparts inconspicuous | Mouthparts well developed, forming a long or short proboscis |
| Thorax without halteres | Thorax with halteres 
|
5 Example Specimens
I'm not fully confident in the ID on this one. I couldn't see for sure that there are two equal pads under the tarsal claws instead of three, but it seems that way. The antennae are 4-segmented, although I'm not sure if the last segment counts as a "segment" rather than a style@ or arista sticking off the third segment. It has three ocelli. The key characteristics generally point to Empididae except "Vein R4+5 usually branched." That vein certainly isn't branched in this specimen, but the key does say "usually."
| |
| Ephemeroptera | Diptera |
Adapted from Merritt R.W., Cummins, K.W., and Berg, M.B. (2019)
The current couplet is highlighted with darker colors and a icon, and couplets leading to this point have a icon.
Leads to Couplet 2:
- Forewings leathery (semiopaque) or hard, at least in the basal half
Couplet 2
Leads to Couplet 4:
- Wings entirely membraneous, usually transparent between veins, but may be variously colored, darkened, or covered with hairs or scales
Couplet 4
Leads to Orthoptera:
- Forewings leathery, with distinct veins
- Femora of hind legs greatly enlarged, suited for jumping
Leads to Coleoptera:
- Forewings hard (called elytra), with veins indistinct; hindwings membraneous but often concealed
- Hind legs suited for walking or swimming
Leads to Couplet 5:
- One pair of wings
Couplet 5
Leads to Couplet 6:
- Two pairs of wings, although hindwings may be small or concealed beneath forewings when wings are folded
Couplet 6
Couplet 5 (You are here)
Leads to Ephemeroptera:
- Two or three tails
- Mouthparts inconspicuous
- Thorax without halteres
Leads to Couplet 7:
- Wings covered with scales or hairs, obscuring venation
Couplet 7
Leads to Couplet 8:
- Wings bare or with minute hairs; venation clearly visible
Couplet 8
Leads to Lepidoptera:
- Wings covered with scales (broadly flattened hairs)
- Mouthparts usually fitted with a coiled sucking tube (proboscis)
Leads to Trichoptera:
- Wings covered with hairs
- Mouthparts without a coiled sucking tube
Leads to Couplet 9:
- Antennae bristle-like and inconspicuous, usually shorter than head
Couplet 9
Leads to Couplet 10:
- Antennae of various shapes, conspicuous, and not bristle-like, and usually much longer than head
Couplet 10
Leads to Ephemeroptera:
- 2 or 3 tails
- Hind wings much smaller than forewings
Leads to Couplet 13:
- No tails
- Wings about equal in size
Couplet 13
Leads to Plecoptera:
- Tarsi 2- or 3-segmented
- 2 conspicuous tails (except reduced in some Nemouridae)
Leads to Couplet 11:
- Tarsi 5-segmented (except a few Hymenoptera have 3 or 4 segments)
- No tails
Couplet 11
Leads to Couplet 12:
- Abdomen broadly joined to thorax
- Front margin of forewing in basal half with many small veins perpendicular to edge
- Wings with more than 20 closed cells
Couplet 12
Leads to Hymenoptera:
- Abdomen with narrow constriction at junction with the thorax
- Marginal veins in basal half of forewing parallel to leading edge
- Wings with fewer than 20 closed cells
- Aquatic forms are very small, usually less than 3 mm long
Leads to Megaloptera:
- Hind wings pleated lengthwise, folded fan-like when closed
- Veins along back edge of wings usually with Y-shaped branches
Leads to Neuroptera:
- Hind wings not pleated, similar to forewings when closed
- Veins along back edge of wings forming many Y-shaped branches
Neuroptera
Leads to Odonata-Zygoptera:
- Wings held together above abdomen or only partly spread when insect is perched
- Eyes separated by more than the width of one eye
- Fore and hind wings similar in size and shape, each with a quadrangle instead of a triangle and subtriangle
- Males with 2 paraprocts
- Females with a fully-developed ovipositor, bearing styli
Leads to Odonata-Anisoptera:
- Wings held horizontally when insect is perched
- Eyes meeting middorsally or separated by a space less than the width of one eye
- Fore and hind wings dissimilar in size and shape, with hind wing considerably wider at base than forewing; each with a triangle and subtriangle
- Males with an epiproct which is either entire or bifid
- Females with or without an ovipositor
Start a Discussion of this Couplet
References
- Merritt R.W., Cummins, K.W., and Berg, M.B. 2019. An Introduction to the Aquatic Insects of North America (Fifth Edition). Kendall/Hunt Publishing Company.
Orders excluded
These orders of Insecta are not included in this key: Psocodea
Description of this key
This key allows identification to order of adult aquatic and semiaquatic insects. Be aware that some invertebrates you might find (including crustaceans, springtails (Collembola), and worms) are not included here. Adult aquatic insects generally have wings present, usually fully developed, conspicuous, and movable, and legs present.