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Artistic view of a Male Pteronarcys californica (Pteronarcyidae) (Giant Salmonfly) Stonefly Adult from the Gallatin River in Montana
Salmonflies
Pteronarcys californica

The giant Salmonflies of the Western mountains are legendary for their proclivity to elicit consistent dry-fly action and ferocious strikes.

Dorsal view of a Grammotaulius betteni (Limnephilidae) (Northern Caddisfly) Caddisfly Larva from the Yakima River in Washington
This is a striking caddis larva with an interesting color pattern on the head. Here are some characteristics I was able to see under the microscope, but could not easily expose for a picture:
- The prosternal horn is present.
- The mandible is clearly toothed, not formed into a uniform scraper blade.
- The seems to be only 2 major setae on the ventral edge of the hind femur.
- Chloride epithelia seem to be absent from the dorsal side of any abdominal segments.
Based on these characteristics and the ones more easily visible from the pictures, this seems to be Grammotaulius. The key's description of the case is spot-on: "Case cylindrical, made of longitudinally arranged sedge or similar leaves," as is the description of the markings on the head, "Dorsum of head light brownish yellow with numerous discrete, small, dark spots." The spot pattern on the head is a very good match to figure 19.312 of Merritt R.W., Cummins, K.W., and Berg, M.B. (2019). The species ID is based on Grammotaulius betteni being the only species of this genus known in Washington state.
27" brown trout, my largest ever. It was the sub-dominant fish in its pool. After this, I hooked the bigger one, but I couldn't land it.
Troutnut is a project started in 2003 by salmonid ecologist Jason "Troutnut" Neuswanger to help anglers and fly tyers unabashedly embrace the entomological side of the sport. Learn more about Troutnut or support the project for an enhanced experience here.

Male Rhithrogena hageni (Western Black Quill) Mayfly Spinner Pictures

Although I could not find the preserved specimen to examine under my good new microscope, I'm tentatively calling it one Rhithrogena hageni, based on apparent similarity to this specimen, which I was able to positively ID.

The relative angle of the penes is a bit shallower in this specimen, but I photographed another specimen from the same collecting trip (and I think even the same swarm, although I don't recall for sure) as the other one, and it had the shallower angle seen on this specimen. I'm guessing it's just variation within the species.

Lateral view of a Male Rhithrogena hageni (Heptageniidae) (Western Black Quill) Mayfly Spinner from the Ruby River in Montana
Male Rhithrogena hageni (Heptageniidae) (Western Black Quill) Mayfly Spinner from the Ruby River in Montana
Male Rhithrogena hageni (Heptageniidae) (Western Black Quill) Mayfly Spinner from the Ruby River in Montana
Male Rhithrogena hageni (Heptageniidae) (Western Black Quill) Mayfly Spinner from the Ruby River in Montana
Male Rhithrogena hageni (Heptageniidae) (Western Black Quill) Mayfly Spinner from the Ruby River in Montana
Dorsal view of a Male Rhithrogena hageni (Heptageniidae) (Western Black Quill) Mayfly Spinner from the Ruby River in Montana
Male Rhithrogena hageni (Heptageniidae) (Western Black Quill) Mayfly Spinner from the Ruby River in Montana
Male Rhithrogena hageni (Heptageniidae) (Western Black Quill) Mayfly Spinner from the Ruby River in Montana
Male Rhithrogena hageni (Heptageniidae) (Western Black Quill) Mayfly Spinner from the Ruby River in Montana
Male Rhithrogena hageni (Heptageniidae) (Western Black Quill) Mayfly Spinner from the Ruby River in Montana
Ventral view of a Male Rhithrogena hageni (Heptageniidae) (Western Black Quill) Mayfly Spinner from the Ruby River in Montana
Male Rhithrogena hageni (Heptageniidae) (Western Black Quill) Mayfly Spinner from the Ruby River in Montana
Male Rhithrogena hageni (Heptageniidae) (Western Black Quill) Mayfly Spinner from the Ruby River in Montana
Ruler view of a Male Rhithrogena hageni (Heptageniidae) (Western Black Quill) Mayfly Spinner from the Ruby River in Montana The smallest ruler marks are 1 mm.

This mayfly was collected from the Ruby River in Montana on July 8th, 2019 and added to Troutnut.com by Troutnut on July 18th, 2019.

Discussions of this Spinner

Puzzled on this mayfly, not even confident on family
1 replies
Posted by Troutnut on Jul 22, 2019
Last reply on Jul 23, 2019 by Millcreek
This seems to be some sort of Heptageniid, based on the two tails and five segments on the hind tarsus. Following the key in Merritt & Cummins seems to lead to Rhithrogena, but it doesn't fit a characteristic described in Needham's Biology of Mayflies for that genus: "Tibia of hind leg in both sexes about 1/4 length of femur."

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Male Rhithrogena hageni (Western Black Quill) Mayfly Spinner Pictures

Collection details
Location: Ruby River, Montana
Date: July 8th, 2019
Added to site: July 18th, 2019
Author: Troutnut
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