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Lateral view of a Female Hexagenia limbata (Ephemeridae) (Hex) Mayfly Dun from the Namekagon River in Wisconsin
Hex Mayflies
Hexagenia limbata

The famous nocturnal Hex hatch of the Midwest (and a few other lucky locations) stirs to the surface mythically large brown trout that only touch streamers for the rest of the year.

Dorsal view of a Grammotaulius betteni (Limnephilidae) (Northern Caddisfly) Caddisfly Larva from the Yakima River in Washington
This is a striking caddis larva with an interesting color pattern on the head. Here are some characteristics I was able to see under the microscope, but could not easily expose for a picture:
- The prosternal horn is present.
- The mandible is clearly toothed, not formed into a uniform scraper blade.
- The seems to be only 2 major setae on the ventral edge of the hind femur.
- Chloride epithelia seem to be absent from the dorsal side of any abdominal segments.
Based on these characteristics and the ones more easily visible from the pictures, this seems to be Grammotaulius. The key's description of the case is spot-on: "Case cylindrical, made of longitudinally arranged sedge or similar leaves," as is the description of the markings on the head, "Dorsum of head light brownish yellow with numerous discrete, small, dark spots." The spot pattern on the head is a very good match to figure 19.312 of Merritt R.W., Cummins, K.W., and Berg, M.B. (2019). The species ID is based on Grammotaulius betteni being the only species of this genus known in Washington state.
27" brown trout, my largest ever. It was the sub-dominant fish in its pool. After this, I hooked the bigger one, but I couldn't land it.
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Male Rhithrogena virilis Mayfly Spinner Pictures

I'm fairly sure this is a specimen of Rhithrogena virilis based on closeup examination of the reproductive anatomy under the microscope (not shown in photos). The other other species of Rhithrogena this large is Rhithrogena flavianula, but the key in Needham's Biology of Mayflies mentions annulation in the abdomen (visible in some images on bugguide.net) more distinct than that on this specimen.

The body and front wing were both about 15.5 mm long, while the cerci were 40 mm long.

There's a pale amber tinge to the anterior areas of the wings, which doesn't show up all that well against the blue background.

Lateral view of a Male Rhithrogena virilis (Heptageniidae) Mayfly Spinner from the South Fork Sauk River in Washington
Male Rhithrogena virilis (Heptageniidae) Mayfly Spinner from the South Fork Sauk River in Washington
Male Rhithrogena virilis (Heptageniidae) Mayfly Spinner from the South Fork Sauk River in Washington
Male Rhithrogena virilis (Heptageniidae) Mayfly Spinner from the South Fork Sauk River in Washington
Dorsal view of a Male Rhithrogena virilis (Heptageniidae) Mayfly Spinner from the South Fork Sauk River in Washington
Male Rhithrogena virilis (Heptageniidae) Mayfly Spinner from the South Fork Sauk River in Washington
Male Rhithrogena virilis (Heptageniidae) Mayfly Spinner from the South Fork Sauk River in Washington
Male Rhithrogena virilis (Heptageniidae) Mayfly Spinner from the South Fork Sauk River in Washington
Male Rhithrogena virilis (Heptageniidae) Mayfly Spinner from the South Fork Sauk River in Washington
Tick marks are 1/16". The body and front wing were both about 15.5 mm long.

Ruler view of a Male Rhithrogena virilis (Heptageniidae) Mayfly Spinner from the South Fork Sauk River in Washington The smallest ruler marks are 1/16".
Male Rhithrogena virilis (Heptageniidae) Mayfly Spinner from the South Fork Sauk River in Washington
Ventral view of a Male Rhithrogena virilis (Heptageniidae) Mayfly Spinner from the South Fork Sauk River in Washington

This mayfly was collected from the South Fork Sauk River in Washington on July 5th, 2017 and added to Troutnut.com by Troutnut on July 6th, 2017.


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Male Rhithrogena virilis Mayfly Spinner Pictures

Collection details
Location: South Fork Sauk River, Washington
Date: July 5th, 2017
Added to site: July 6th, 2017
Author: Troutnut
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