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Lateral view of a Male Baetis (Baetidae) (Blue-Winged Olive) Mayfly Dun from Mystery Creek #43 in New York
Blue-winged Olives
Baetis

Tiny Baetis mayflies are perhaps the most commonly encountered and imitated by anglers on all American trout streams due to their great abundance, widespread distribution, and trout-friendly emergence habits.

Case view of a Pycnopsyche guttifera (Limnephilidae) (Great Autumn Brown Sedge) Caddisfly Larva from the Yakima River in Washington
It's only barely visible in one of my pictures, but I confirmed under the microscope that this one has a prosternal horn and the antennae are mid-way between the eyes and front of the head capsule.

I'm calling this one Pycnopsyche, but it's a bit perplexing. It seems to key definitively to at least Couplet 8 of the Key to Genera of Limnephilidae Larvae. That narrows it down to three genera, and the case seems wrong for the other two. The case looks right for Pycnopsyche, and it fits one of the key characteristics: "Abdominal sternum II without chloride epithelium and abdominal segment IX with only single seta on each side of dorsal sclerite." However, the characteristic "metanotal sa1 sclerites not fused, although often contiguous" does not seem to fit well. Those sclerites sure look fused to me, although I can make out a thin groove in the touching halves in the anterior half under the microscope. Perhaps this is a regional variation.

The only species of Pycnopsyche documented in Washington state is Pycnopsyche guttifera, and the colors and markings around the head of this specimen seem to match very well a specimen of that species from Massachusetts on Bugguide. So I am placing it in that species for now.

Whatever species this is, I photographed another specimen of seemingly the same species from the same spot a couple months later.
27" brown trout, my largest ever. It was the sub-dominant fish in its pool. After this, I hooked the bigger one, but I couldn't land it.
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Dun Variants

Like most common names,"Dun Variant" can refer to more than one taxon. They're previewed below, along with 4 specimens. For more detail click through to the scientific names.

Mayfly Species Isonychia bicolor

These are often called Dun Variants.
This is by far the most important species of Isonychia. Many angling books once split its credit with the species Isonychia sadleri and Isonychia harperi, but entomologists have since discovered that those are just variations of this abundant species.

See the main Isonychia page for more about these intriguing mayflies.
Lateral view of a Female Isonychia bicolor (Isonychiidae) (Mahogany Dun) Mayfly Dun from the Namekagon River in Wisconsin
Lateral view of a Female Isonychia bicolor (Isonychiidae) (Mahogany Dun) Mayfly Spinner from the West Branch of Owego Creek in New York
I collected this female together with a male.
Dorsal view of a Isonychia bicolor (Isonychiidae) (Mahogany Dun) Mayfly Nymph from the Beaverkill River in New York

Mayfly Species Attenella attenuata

These are very rarely called Dun Variants.
This intriguing species has received a lot of attention in past angling books. Recent authors suspect that much of this credit was a case of mistaken identity, with Attenella attenuata receiving praise for the hatches of Drunella lata and Dannella simplex. Much of the credit was legitimate and accurate, but this species is no longer thought to be on par with its most popular cousins in Ephemerella and Drunella.

I have several specimens listed under this species, but I'm not positive the identification is correct.
Lateral view of a Female Attenella attenuata (Ephemerellidae) (Blue-Winged Olive) Mayfly Dun from the Namekagon River in Wisconsin
This specimen came from the same hatch as a male.

Dun Variants

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