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Lateral view of a Female Hexagenia limbata (Ephemeridae) (Hex) Mayfly Dun from the Namekagon River in Wisconsin
Hex Mayflies
Hexagenia limbata

The famous nocturnal Hex hatch of the Midwest (and a few other lucky locations) stirs to the surface mythically large brown trout that only touch streamers for the rest of the year.

Dorsal view of a Pycnopsyche guttifera (Limnephilidae) (Great Autumn Brown Sedge) Caddisfly Larva from the Yakima River in Washington
This specimen appears to be of the same species as this one collected in the same spot two months earlier. The identification of both is tentative. This one suffered some physical damage before being photographed, too, so the colors aren't totally natural. I was mostly photographing it to test out some new camera setting idea, which worked really well for a couple of closeups.
27" brown trout, my largest ever. It was the sub-dominant fish in its pool. After this, I hooked the bigger one, but I couldn't land it.
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Western Ginger Quills

Like most common names,"Western Ginger Quill" can refer to more than one taxon. They're previewed below, along with 5 specimens. For more detail click through to the scientific names.

Mayfly Genus Ecdyonurus

These are often called Western Ginger Quills.
This fairly new genus holds some Western species that used to be in Nixe and were in Heptagenia before that. They are mentioned in books by those generic names. Ecdyonurus simpliciodes is especially important.
Lateral view of a Male Ecdyonurus criddlei (Heptageniidae) (Little Slate-Winged Dun) Mayfly Spinner from the Bitterroot River in Montana
This one was very puzzling to identify. It's clearly Heptageniidae. It keys pretty easily to couplet 58 in Merritt, Cummins, & Berg 5th Edition, but that's where things get tricky.

58: Male penes distinctly L-shaped; Segment 1 of foretarsi usually 1/3 to 2/3 the length of segment 2
58': Males penes not distinctly L-shaped; Segment 1 of foretarsi usually 1/5 to 1/2 the length of segment 2

The characteristics conflict here, as I'd say the penes ARE L-shaped, but the fore tarsal ratio is less than 1/3. Maybe the weasel word "usually" allows for that, and it's some kind of Stenonema, for which I'm not aware of any records in Montana, let alone western Montana.

Going by the tarsal ratio and assuming it's not Stenonema, everything in couplet 60 points to Afghanurus: contiguous compound eyes, genitalia similar to fig 13.222 in MC&B, and weakly developed basal costal crossveins in the forewings. But I looked up the original descriptions of the three Afghanurus species, and the body sizes (flowersi 5-7 mm, inconspicua 4 mm, joernensis 6-7 mm) are way below this one's 10 mm.

This led me to notice that the closely related Ecdyonurus is not included in the key at all, so I checked the descriptions of those species and found that Ecdyonurus criddlei fits very well: the color of the legs, the markings on the tergites and sternites, the short fore tarsus and ratios of the segments, etc. It was originally described as 7 mm, smaller than the 10 mm body of this specimen, but synonyms have been described with larger sizes including wing lengths up to 11 mm for females for Heptagenia salvini.

This specimen was found with a female nearby, which I think is probably the same species, although it is difficult to be certain.
Ecdyonurus criddlei (Heptageniidae) (Little Slate-Winged Dun) Mayfly Nymph from Kintla Lake in Montana
This is an interesting Heptageniid mayfly since in western Montana it is only found in cold lakes, especially those in Glacier National Park.

Mayfly Species Cinygmula reticulata

These are sometimes called Western Ginger Quills.
Cinygmula reticulata is probably the second most important species of Cinygmula behind Cinygmula ramaleyi, perhaps because the waters where it can be found in good numbers are often more remote. They have been reported as abundant in many high country streams of the Southern Rockies as well as the High Sierra's Eastern slope. An obvious difference in their coloration may be the easiest way to tell them apart. Cinygmula ramaleyi is more somber with a brownish body and dark gray wings and is often confused with the similar sized and colored Ephemerella tibialis, in spite of the difference in tail counts. Cinygmula reticulata on the other hand is a bright cinnamon dorsally with pale creamy legs and pale wings that are often a brilliant canary yellow. This is one of North America's most beautiful mayflies.
Female Cinygmula reticulata (Heptageniidae) (Western Ginger Quill) Mayfly Dun from the Touchet River in Washington
Lateral view of a Male Cinygmula reticulata (Heptageniidae) (Western Ginger Quill) Mayfly Spinner from Mystery Creek #237 in Montana
The lengths of the wing and body, measured with a caliper, are both 8 mm.

Keys in Needham's 1935 Biology of Mayflies point to either Cinygmula reticulata or Cinygmula gartrelli. It seems to have crossveins in costal half of forewing only, slightly margined with brown and wings tinged with amber at base and along costal margin of both wing (gartrelli) as opposed to all crossveins of both wings faintly but broadly margined with pale smoky and wings entirely amber-tinged (although there is a slight amber tinge throughout, just more pronounced in places) as in reticulata. However, wing length reported for reticulata (9 mm) is closer to this specimen than gartrelli (10 mm). Ventral median marks are supposed to be "traces" for reticulata and "present" for gartrelli. Descriptions for both species involve semi-hyaline anterior abdominal segments not present on my specimens. Distribution records suggest reticulata lives nearby, so I'm going with that, but I can't confidently rule out gartrelli.
Cinygmula reticulata (Heptageniidae) (Western Ginger Quill) Mayfly Nymph from the Big Thompson River in Montana
I collected several live specimens of nymphs and reared them to the imago stage. They were C. reticulata. The interesting thing is they were collected in May and were emerging along with Rhithrogena (March Brown). This seems to be an overlooked hatch since in some rivers it emerges very early, before runoff.

Mayfly Species Ecdyonurus simplicioides

These are sometimes called Western Ginger Quills.
This is an important Western hatch.

Western Ginger Quills

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