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Lateral view of a Female Hexagenia limbata (Ephemeridae) (Hex) Mayfly Dun from the Namekagon River in Wisconsin
Hex Mayflies
Hexagenia limbata

The famous nocturnal Hex hatch of the Midwest (and a few other lucky locations) stirs to the surface mythically large brown trout that only touch streamers for the rest of the year.

Dorsal view of a Grammotaulius betteni (Limnephilidae) (Northern Caddisfly) Caddisfly Larva from the Yakima River in Washington
This is a striking caddis larva with an interesting color pattern on the head. Here are some characteristics I was able to see under the microscope, but could not easily expose for a picture:
- The prosternal horn is present.
- The mandible is clearly toothed, not formed into a uniform scraper blade.
- The seems to be only 2 major setae on the ventral edge of the hind femur.
- Chloride epithelia seem to be absent from the dorsal side of any abdominal segments.
Based on these characteristics and the ones more easily visible from the pictures, this seems to be Grammotaulius. The key's description of the case is spot-on: "Case cylindrical, made of longitudinally arranged sedge or similar leaves," as is the description of the markings on the head, "Dorsum of head light brownish yellow with numerous discrete, small, dark spots." The spot pattern on the head is a very good match to figure 19.312 of Merritt R.W., Cummins, K.W., and Berg, M.B. (2019). The species ID is based on Grammotaulius betteni being the only species of this genus known in Washington state.
27" brown trout, my largest ever. It was the sub-dominant fish in its pool. After this, I hooked the bigger one, but I couldn't land it.
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Mayfly Species Eurylophella lutulenta (Chocolate Duns)

The only account of this species I've found in angling literature is in the Leonards' Mayflies of Michigan Trout Streams. They remark that it is the size of Ephemerella subvaria, much larger than any Eurylophella nymphs I've collected, and its color is a "rather bright yellowish-brown."

Where & when

Time of year : Late May through early June

Preferred waters: Lakes and large rivers

In 4 records from GBIF, adults of this species have been collected during June (75%) and April (25%).

In 1 record from GBIF, this species has been collected at elevation of 2723 ft.

Species Range

Physical description

Most physical descriptions on Troutnut are direct or slightly edited quotes from the original scientific sources describing or updating the species, although there may be errors in copying them to this website. Such descriptions aren't always definitive, because species often turn out to be more variable than the original describers observed. In some cases, only a single specimen was described! However, they are useful starting points.

Male Spinner

Described in Needham et al (1935) as Ephemerella lutulenta
Body length: 8-9 mm
Wing length: 10 mm

A large species of the bicolor group (now a synonym of Eurylophella bicolor); imago distinguished by the sprinkling of very small dark dots on legs, thorax and abdomen.

Face dark brown; a reddish grey streak on the median carina, and two lateral streaks to bases of antennae. Thorax dark reddish brown. Legs greenish yellow, claws brown. Wings hyaline; venation hyaline. Abdomen blackish brown dorsally, segments 8 and 9 rather paler in color. Posterior margins of tergites narrowly darker reddish brown. Pleural fold blackish brown on the margin; a black streak in each stigmatic region. Legs, venter of thorax and the entire abdomen sprinkled with very fine dark dots. Tails reddish brown near the base, paler distally; joinings dark brown.

Nymph

Described in Needham et al (1935) as Ephemerella lutulenta

In the male nymph the occipital tubercles are practically wanting, but are much better developed in the female. Rows of dorsal spines diverge to rearward. The spines on tergites 1-3 are long, erect and rather finger-like, and bend backward; on 8 and 9 they are almost obsolete. The postero-lateral spines are well developed, the spine on segment 3 being fully twice as long as its width at the base; the spines on segments 4-8 are very long and curved, bearing spinules and hairs on the margins. The operculum is veined longitudinally and usually bears several to many small pale dots. The color varies from dirty brown to a dark blackish brown; legs brown with pale bands, the femora with pale spots. Tails rather unicolorous brown, the cross bands of paler color less distinct than in E. temporalis (now a synonym of Eurylophella temporalis). An inhabitant of rather large bodies of water.


Start a Discussion of Eurylophella lutulenta

References

Mayfly Species Eurylophella lutulenta (Chocolate Duns)

Species Range
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