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Lateral view of a Female Hexagenia limbata (Ephemeridae) (Hex) Mayfly Dun from the Namekagon River in Wisconsin
Hex Mayflies
Hexagenia limbata

The famous nocturnal Hex hatch of the Midwest (and a few other lucky locations) stirs to the surface mythically large brown trout that only touch streamers for the rest of the year.

Dorsal view of a Ephemerella mucronata (Ephemerellidae) Mayfly Nymph from the Yakima River in Washington
This is an interesting one. Following the keys in Merritt R.W., Cummins, K.W., and Berg, M.B. (2019) and Jacobus et al. (2014), it keys clearly to Ephemerella. Jacobus et al provide a key to species, but some of the characteristics are tricky to interpret without illustrations. If I didn't make any mistakes, this one keys to Ephemerella mucronata, which has not previously been reported any closer to here than Montana and Alberta. The main character seems to fit well: "Abdominal terga with prominent, paired, subparallel, spiculate ridges." Several illustrations or descriptions of this holarctic species from the US and Europe seem to match, including the body length, tarsal claws and denticles, labial palp, and gill shapes. These sources include including Richard Allen's original description of this species in North America under the now-defunct name E. moffatae in Allen RK (1977) and the figures in this description of the species in Italy.
27" brown trout, my largest ever. It was the sub-dominant fish in its pool. After this, I hooked the bigger one, but I couldn't land it.
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Alderfly Genus Sialis

Alderflies are typically mistaken for small hellgrammites as larvae and large caddisflies as adults. They turn up often enough in nymph samples that it's nice to know what they look like. There are rumors of their occasional importance to trout, but the average anglers is not likely to encounter it.

Where & when

Time of year : Early season

In 294 records from GBIF, adults of this genus have mostly been collected during April (35%), May (27%), June (25%), and July (8%).

In 85 records from GBIF, this genus has been collected at elevations ranging from 495 to 11526 ft, with an average (median) of 3182 ft.

Genus Range

Hatching behavior

Time of day : Evening

Pupae burrow into the ground near to, but not in, the stream. Because the emergence takes place on land it is unimportant to trout.

Egg-Laying behavior

Time of day: Daytime

The mating adults gather over the water in good numbers but do not usually fall into the water unless wind knocks them off course.

They lay their eggs on dry objects near the water. Ernest Schwiebert describes their egg placement in Matching the Hatch:

The egg masses are deposited on rocks, limbs, leaves and bridges where they are over running water and in the sunlight. They are placed so that the hatching larvae fall into the stream.

Larva & pupa biology

Current speed: Slow

Substrate: Silt

The larvae are burrowers like the Ephemeridae mayfly nymphs, digging down into the silt.

Specimens of the Alderfly Genus Sialis

1 Male Adult
4 Larvae

Start a Discussion of Sialis

References

Alderfly Genus Sialis

Taxonomy
Species in Sialis
Sialis hamata
1
2
Species in Sialis: Sialis hamata
23 species (Sialis aequalis, Sialis americana, Sialis arvalis, Sialis bilobata, Sialis californica, Sialis concava, Sialis contigua, Sialis cornuta, Sialis dreisbachi, Sialis glabella, Sialis hasta, Sialis infumata, Sialis iola, Sialis itasca, Sialis joppa, Sialis mohri, Sialis nevadensis, Sialis nina, Sialis occidens, Sialis rotunda, Sialis spangleri, Sialis vagans, and Sialis velata) aren't included.
Genus Range
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