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Lateral view of a Female Hexagenia limbata (Ephemeridae) (Hex) Mayfly Dun from the Namekagon River in Wisconsin
Hex Mayflies
Hexagenia limbata

The famous nocturnal Hex hatch of the Midwest (and a few other lucky locations) stirs to the surface mythically large brown trout that only touch streamers for the rest of the year.

Dorsal view of a Setvena wahkeena (Perlodidae) (Wahkeena Springfly) Stonefly Nymph from Mystery Creek #199 in Washington
As far as I can tell, this species has only previously been reported from one site in Oregon along the Columbia gorge. However, the key characteristics are fairly unmistakable in all except for one minor detail:
— 4 small yellow spots on frons visible in photos
— Narrow occipital spinule row curves forward (but doesn’t quite meet on stem of ecdysial suture, as it's supposed to in this species)
— Short spinules on anterior margin of front legs
— Short rposterior row of blunt spinules on abdominal tergae, rather than elongated spinules dorsally
I caught several of these mature nymphs in the fishless, tiny headwaters of a creek high in the Wenatchee Mountains.
27" brown trout, my largest ever. It was the sub-dominant fish in its pool. After this, I hooked the bigger one, but I couldn't land it.
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Caddisfly Species Dolophilodes distincta (Tiny Black Gold Speckled-Winged Caddisflies)

What a weird species is Dolophilodes distinctus. It emerges year-round, and the female adults in the winter do not have wings. In the summer they do, and in the spring there is a mix.

This is the most important species of Dolophilodes in the East and Midwest. It is never an intense emergence, but it may be present at times of the year when nothing else is available.

Where & when

Time of year : All year; typically two peaks in winter and again in midsummer

Species Range

Hatching behavior

Time of day : Morning and afternoon

The pupae swim to the surface and then scoot across it to the shore to emerge. Swisher and Richards comment on the importance of this behavior in Selective Trout:

When fishing to an emergence, the adult and emerger imitations are usueless. The angler must use a hackled pupa skated over the water toward shore.

Larva & pupa biology

Environmental tolerance: Prefers cold water

Specimens of the Caddisfly Species Dolophilodes distincta

1 Male Adult
1 Female Adult
1 Larva

Discussions of Dolophilodes distincta


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References

Caddisfly Species Dolophilodes distincta (Tiny Black Gold Speckled-Winged Caddisflies)

Taxonomy
Species Range
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