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Artistic view of a Male Pteronarcys californica (Pteronarcyidae) (Giant Salmonfly) Stonefly Adult from the Gallatin River in Montana
Salmonflies
Pteronarcys californica

The giant Salmonflies of the Western mountains are legendary for their proclivity to elicit consistent dry-fly action and ferocious strikes.

Dorsal view of a Setvena wahkeena (Perlodidae) (Wahkeena Springfly) Stonefly Nymph from Mystery Creek #199 in Washington
As far as I can tell, this species has only previously been reported from one site in Oregon along the Columbia gorge. However, the key characteristics are fairly unmistakable in all except for one minor detail:
— 4 small yellow spots on frons visible in photos
— Narrow occipital spinule row curves forward (but doesn’t quite meet on stem of ecdysial suture, as it's supposed to in this species)
— Short spinules on anterior margin of front legs
— Short rposterior row of blunt spinules on abdominal tergae, rather than elongated spinules dorsally
I caught several of these mature nymphs in the fishless, tiny headwaters of a creek high in the Wenatchee Mountains.
27" brown trout, my largest ever. It was the sub-dominant fish in its pool. After this, I hooked the bigger one, but I couldn't land it.
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Identification: Key to Genera of Ephemeridae Duns and Spinners, Couplet 2

Identification: Key to Genera of Ephemeridae Duns and Spinners, Couplet 2

Adapted from Merritt R.W., Cummins, K.W., and Berg, M.B. (2019)
This couplet refers figures (usually designated 'sf') from this source.
You will need the source (make sure to get the same edition!) to see them.
Option 1Option 2
Head with frons greatly expanded below compound eyes (source fig 13.251)Head with frons not extending below compound eyes (source fig 13.252)
Wing membranes tinted amber or purplish brownWing membranes in spinners transparent or occasionally slightly tinted yellow
Male penes recurved ventrally (source fig 13.250)Male penes not recurved as described, but may be hooked or curved medially (source fig 13.253)
3 Example Specimens
5 Example Specimens
Litobrancha Hexagenia
The current couplet is highlighted with darker colors and a icon, and couplets leading to this point have a icon.
Leads to Ephemera:
  • Three tails (i.e., terminal filament subequal to cerci)
  • Forewings, and sometimes hind wings, prominently mottled with dark markings
  • Crossveins of forewings near bullae crowded together (source fig 13.168a)
Leads to Couplet 2:
  • Two main tails (i.e., terminal filament vestigial or clearly shorter than cerci)
  • Forewings and hingwings not prominently mottled, but may be shaded along margins
  • Crossveins of forewings near bullae not crowded together
Couplet 2
Couplet 2

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Couplet 2 (You are here)
Leads to Litobrancha
(Litobrancha recurvata):
  • Head with frons greatly expanded below compound eyes (source fig 13.251)
  • Wing membranes tinted amber or purplish brown
  • Male penes recurved ventrally (source fig 13.250)
Leads to Hexagenia:
  • Head with frons not extending below compound eyes (source fig 13.252)
  • Wing membranes in spinners transparent or occasionally slightly tinted yellow
  • Male penes not recurved as described, but may be hooked or curved medially (source fig 13.253)

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References

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