Artistic view of a Male Pteronarcys californica (Pteronarcyidae) (Giant Salmonfly) Stonefly Adult from the Gallatin River in Montana

Salmonflies
Pteronarcys californica

The giant Salmonflies of the Western mountains are legendary for their proclivity to elicit consistent dry-fly action and ferocious strikes.

For how many years, I wonder, have these huge slabs of rock sheltered wild brook trout?

From Mystery Creek # 89 in New York

Stream photos

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Grammotaulius betteni (Northern Caddisfly) Caddisfly Larva Grammotaulius betteni (Northern Caddisfly) Caddisfly Larva

Dorsal view of a Grammotaulius betteni (Limnephilidae) (Northern Caddisfly) Caddisfly Larva from the Yakima River in Washington

This is a striking caddis larva with an interesting color pattern on the head. Here are some characteristics I was able to see under the microscope, but could not easily expose for a picture:
- The prosternal horn is present.
- The mandible is clearly toothed, not formed into a uniform scraper blade.
- The seems to be only 2 major setae on the ventral edge of the hind femur.
- Chloride epithelia seem to be absent from the dorsal side of any abdominal segments.
Based on these characteristics and the ones more easily visible from the pictures, this seems to be Grammotaulius. The key's description of the case is spot-on: "Case cylindrical, made of longitudinally arranged sedge or similar leaves," as is the description of the markings on the head, "Dorsum of head light brownish yellow with numerous discrete, small, dark spots." The spot pattern on the head is a very good match to figure 19.312 of Merritt R.W., Cummins, K.W., and Berg, M.B. (2019). The species ID is based on Grammotaulius betteni being the only species of this genus known in Washington state.

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27" brown trout, my largest ever. It was the sub-dominant fish in its pool. After this, I hooked the bigger one, but I couldn't land it.

Troutnut is a project started in 2003 by salmonid ecologist Jason "Troutnut" Neuswanger to help anglers and fly tyers unabashedly embrace the entomological side of the sport. Learn more about Troutnut or support the project for an enhanced experience here.

Helicopsyche borealis (Speckled Peter) Caddisfly Pupa Pictures

I'm calling this one very tentatively Helicopsyche borealis, with some big caveats that I might be wrong.

It seems to key fairly easily to Helicopsychidae, except at the last couplet in Merritt & Cummins (5th ed) it's supposed to have “anal processes short and straight, each with several mesal setae and 2 long apical setae,” whereas this one has VERY short, straight processes with 3 long apical setae and no mesal ones.

Additionally, the only species of Helicopsychidae documented in this region is Helicopsyche borealis, which is supposed to emerge much later in the summer.