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Artistic view of a Male Pteronarcys californica (Pteronarcyidae) (Giant Salmonfly) Stonefly Adult from the Gallatin River in Montana
Salmonflies
Pteronarcys californica

The giant Salmonflies of the Western mountains are legendary for their proclivity to elicit consistent dry-fly action and ferocious strikes.

Case view of a Pycnopsyche guttifera (Limnephilidae) (Great Autumn Brown Sedge) Caddisfly Larva from the Yakima River in Washington
It's only barely visible in one of my pictures, but I confirmed under the microscope that this one has a prosternal horn and the antennae are mid-way between the eyes and front of the head capsule.

I'm calling this one Pycnopsyche, but it's a bit perplexing. It seems to key definitively to at least Couplet 8 of the Key to Genera of Limnephilidae Larvae. That narrows it down to three genera, and the case seems wrong for the other two. The case looks right for Pycnopsyche, and it fits one of the key characteristics: "Abdominal sternum II without chloride epithelium and abdominal segment IX with only single seta on each side of dorsal sclerite." However, the characteristic "metanotal sa1 sclerites not fused, although often contiguous" does not seem to fit well. Those sclerites sure look fused to me, although I can make out a thin groove in the touching halves in the anterior half under the microscope. Perhaps this is a regional variation.

The only species of Pycnopsyche documented in Washington state is Pycnopsyche guttifera, and the colors and markings around the head of this specimen seem to match very well a specimen of that species from Massachusetts on Bugguide. So I am placing it in that species for now.

Whatever species this is, I photographed another specimen of seemingly the same species from the same spot a couple months later.
27" brown trout, my largest ever. It was the sub-dominant fish in its pool. After this, I hooked the bigger one, but I couldn't land it.
Troutnut is a project started in 2003 by salmonid ecologist Jason "Troutnut" Neuswanger to help anglers and fly tyers unabashedly embrace the entomological side of the sport. Learn more about Troutnut or support the project for an enhanced experience here.

Mayfly Species Stenonema mediopunctatum (Cream Cahills)

I have seen no mention of this species as a fishable hatch in angling literature, but it is noteworthy for the unusual coloration of the male spinners, which Malcolm and Knopp describe in Mayflies: An Angler's Study of Trout Water Ephemeroptera :

...the male only exhibits this coloration on segments eight to ten, against an otherwise whitish to translucent body.

Where & when

Time of year : Late June to early October, peaking in early August

In 5 records from GBIF, adults of this species have been collected during July (80%) and June (20%).

Species Range

Nymph biology

Current speed: Fast

Substrate: Gravel and rock

Physical description

Most physical descriptions on Troutnut are direct or slightly edited quotes from the original scientific sources describing or updating the species, although there may be errors in copying them to this website. Such descriptions aren't always definitive, because species often turn out to be more variable than the original describers observed. In some cases, only a single specimen was described! However, they are useful starting points.

Male Spinner

Body length: 8-9 mm
Wing length: 8-10 mm

A species of Stenonema pulchellum group; thorax almost wholly deep black brown; spiracular dots usually present; tails not, darker at joinings.

Frontal portion of head pale whitish; vertex sepia brown with ruddy shading next to the eyes. Thorax, dorsally and ventrally, deep black brown, except a whitish patch on pleura anterior to wing roots and slight paler shading below wing roots. Posterior portion of scutellum with faint paler shading; a small whitish median dot anterior to scutellum, and two small white lateral streaks. Legs whitish; median and apical purplish bands on femora; fore tarsus black-tipped at apex; basal joint of fore tarsus more than half as long as second. Wings hyaline; veins fine, black, cross veins in costo-apical region slightly thickened.

Abdominal segments 2-7 hyaline white; scarcely a trace of darker posterior margins, only a small dark transverse dash on posterior margin of each, at mid-dorsal line. Black spiracular dots present on segments 4-7 (may be obsolete). Segments 8-10 opaque, whitish; tergite 8 in posterior half, and all of 9, shaded with light sepia brown. Forceps whitish; genitalia of the pulchellum type(?). Tails wholly white, not darker at joinings.

The very dark thorax, short dark median dash on tergites and fine veins distinguish this species from its near allies.

Described as S. nepotellum subspecies mediopunctatum

Body length 8 mm, wing length 8-10 mm

A species of the Stenonema pulchellum group; spiracular dots present; tails dark-ringed; reddish stain in stigmatic area of fore wing.

Head very similar to S. rubromaculatum (now a synonym of Stenonema modestum). Mesonotum ochre-brown; anterolateral margins narrowly whitish. Scutellum whitish, also lateral margins to base of fore wing. Metanotum largely ochre-brown; median elevation whitish. Pleura yellowish-white, with brown shading as in rubromaculatum; anterior to wing roots shaded with light reddish. Legs very much as in rubromaculatum; posterior margin of fore coxa narrowly black; less reddish shading laterally than in that species; purplish bands on femora also less distinct, may be wanting on hind leg. Wings as in rubromaculatum.

Abdominal segments 1-7 hyaline, pale yellowish white; posterior margins of tergites somewhat more distinctly black than in rubromaculatum; black spiracular dots well defined. Segments 8-10 opaque white; tergites largely suffused with pinkish brown; on anterior margin of 8, slight pale submedian wedge-shaped marks; posterior margin of 10 also white. Genitalia of rubromaculatum type. Tails pale; joinings narrowly brown.

In the imago stage, this species is very close to S. rubromaculatum and S. annexum (now a synonym of Stenonema modestum); may be distinguished as indicated under those species. Ventral markings of nymph are distinct from either of these, and very similar likewise to Stenonema ithaca.

Specimens of the Mayfly Species Stenonema mediopunctatum

2 Nymphs

Discussions of Stenonema mediopunctatum

M. ithaca in M. mediopunctatum section?
3 replies
Posted by GONZO on Sep 1, 2012
Last reply on Sep 4, 2012 by Entoman
Hi Jason,

In this section, the Midwestern nymphs (#573 & #574) with the dark irregular ventral bars across the anterior portion of the sternites look like Maccaffertium mediopunctatum arwini (the Midwestern ssp.), but the two Eastern duns (#733 & #765) and the associated shuck (of #765) and nymph (#764) look more like Maccaffertium ithaca to me.

Three Maccaffertium species can have very similar ventral markings in the nymph—dark, sinuate, chevron-shaped bars on many of the sternites and dark lateral marks (sometimes connected to form an inverted U-shaped mark) on segment 9. The Eastern mediopunctatum subspecies, M. m. mediopunctatum, has these markings, as does M. ithaca. Similar markings also appear as a less-common variant marking of M. modestum (or the M. modestum species complex). However, these species differ in the length and location of posterolateral projections, leg markings, the appearance of the subs and adults, and size.

Although interpretation of posterolateral projections can be tricky, those projections should help to separate the nymph (and husk) from mediopunctatum. On mediopunctatum, projections are usually on segments 3-9, 4-9, or 5-9, and those on 8 and 9 are fairly long. On ithaca, projections are usually on segments 6-9 or 7-9, and those on 8 and 9 are somewhat shorter (when compared to mediopunctatum). The twin brown bands on the femora of the nymphs should also help to separate them from modestum and mediopunctatum (usually three or four in those species).

The brown posterior margins and median dorsal stripes of the duns (similar to those found in M. vicarium) are typical of ithaca. In McDunnough’s original description of mediopunctatum (1926), he mentions that some of his (paratype) specimens were reared from subimagos, and he describes those subimagos as “quite pale whitish in coloration.”

Size might also be somewhat helpful in distinguishing these specimens from M. m. mediopunctatum (about 7-10 mm at maturity) and modestum (about 8-11 mm at maturity). M. ithaca is about 9-14 mm at maturity. The relatively mature nymph (#764) is at least 11 mm, the female dun (#733) is about 13 mm, and the male dun (#765) is about 11 mm.

When all of these factors are considered, it seems to me that M. ithaca is a more likely ID for the Eastern specimens. (See Bednarik and McCafferty 1979 and Lewis 1974.) I would suggest the following placement for specimens currently in this section:

M. ithaca nymph: http://www.troutnut.com/specimen/764
M. ithaca female dun: http://www.troutnut.com/specimen/733
M. ithaca male dun: http://www.troutnut.com/specimen/765
M. mediopunctatum arwini nymph: http://www.troutnut.com/specimen/573
M. mediopunctatum arwini nymph: http://www.troutnut.com/specimen/574

Best,
Lloyd


Start a Discussion of Stenonema mediopunctatum

References

Mayfly Species Stenonema mediopunctatum (Cream Cahills)

Species Range
Common Name
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