Salmonflies
Pteronarcys californica
The giant Salmonflies of the Western mountains are legendary for their proclivity to elicit consistent dry-fly action and ferocious strikes.
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This is a striking caddis larva with an interesting color pattern on the head. Here are some characteristics I was able to see under the microscope, but could not easily expose for a picture:
- The prosternal horn is present.
- The mandible is clearly toothed, not formed into a uniform scraper blade.
- The seems to be only 2 major setae on the ventral edge of the hind femur.
- Chloride epithelia seem to be absent from the dorsal side of any abdominal segments.
Based on these characteristics and the ones more easily visible from the pictures, this seems to be Grammotaulius. The key's description of the case is spot-on: "Case cylindrical, made of longitudinally arranged sedge or similar leaves," as is the description of the markings on the head, "Dorsum of head light brownish yellow with numerous discrete, small, dark spots." The spot pattern on the head is a very good match to figure 19.312 of Merritt R.W., Cummins, K.W., and Berg, M.B. (2019). The species ID is based on Grammotaulius betteni being the only species of this genus known in Washington state.
- The prosternal horn is present.
- The mandible is clearly toothed, not formed into a uniform scraper blade.
- The seems to be only 2 major setae on the ventral edge of the hind femur.
- Chloride epithelia seem to be absent from the dorsal side of any abdominal segments.
Based on these characteristics and the ones more easily visible from the pictures, this seems to be Grammotaulius. The key's description of the case is spot-on: "Case cylindrical, made of longitudinally arranged sedge or similar leaves," as is the description of the markings on the head, "Dorsum of head light brownish yellow with numerous discrete, small, dark spots." The spot pattern on the head is a very good match to figure 19.312 of Merritt R.W., Cummins, K.W., and Berg, M.B. (2019). The species ID is based on Grammotaulius betteni being the only species of this genus known in Washington state.
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White-Winged Curses
This common name refers to only one genus. Click its scientific name to learn more.
Mayfly Genus Tricorythodes
These are very rarely called White-Winged Curses.
A cult following is something to which few insects can lay claim, but the tiny Tricorythodes mayflies certainly qualify. Their widespread, reliable, heavy hatches draw impressive rises of ultra-selective trout which demand the most of a technical dry-fly angler's skills.
It is surprising that such a great hatch took so long to come to the attention of fly fishermen. The Tricos were first introduced to anglers in a 1969 Outdoor Life article by Vincent Marinaro, who misidentified them as Caenis. By the early 1970s the identification had been corrected but Swisher and Richards still wrote in Selective Trout, "Few anglers are familiar with these extremely small but important mayflies." The next wave of publications boosted Tricorythodes to its current fame. I suspect their early dismissal was due in part to tackle limitations; anglers in the 1950s had no means to effectively tie and present size 22-28 flies.
It is surprising that such a great hatch took so long to come to the attention of fly fishermen. The Tricos were first introduced to anglers in a 1969 Outdoor Life article by Vincent Marinaro, who misidentified them as Caenis. By the early 1970s the identification had been corrected but Swisher and Richards still wrote in Selective Trout, "Few anglers are familiar with these extremely small but important mayflies." The next wave of publications boosted Tricorythodes to its current fame. I suspect their early dismissal was due in part to tackle limitations; anglers in the 1950s had no means to effectively tie and present size 22-28 flies.
This female Trico was collected with an associated male and many others.
See 10 more specimens...
References
- Merritt R.W., Cummins, K.W., and Berg, M.B. 2019. An Introduction to the Aquatic Insects of North America (Fifth Edition). Kendall/Hunt Publishing Company.