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Lateral view of a Female Hexagenia limbata (Ephemeridae) (Hex) Mayfly Dun from the Namekagon River in Wisconsin
Hex Mayflies
Hexagenia limbata

The famous nocturnal Hex hatch of the Midwest (and a few other lucky locations) stirs to the surface mythically large brown trout that only touch streamers for the rest of the year.

27" brown trout, my largest ever. It was the sub-dominant fish in its pool. After this, I hooked the bigger one, but I couldn't land it.
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Western Gordon Quills

Like most common names,"Western Gordon Quill" can refer to more than one taxon. They're previewed below, along with 8 specimens. For more detail click through to the scientific names.

Mayfly Genus Epeorus

These are sometimes called Western Gordon Quills.
There is remarkable variety of form and color within this prolific genus of fast-water mayflies. Different species are found across the country, and several cause good hatches. Fly anglers are likely to encounter the lesser species on occasion, too.

The best Epeorus hatch in the East is Epeorus pleuralis, the famous Quill Gordon, the first abundant large mayfly hatch of the year. Epeorus vitreus comes a little later and is important in both the East and Midwest.

In the West, Epeorus longimanus dominates in fast, high-altitude streams, while Epeorus albertae inhabits slower and lower waters.
Artistic view of a Male Epeorus deceptivus (Heptageniidae) Mayfly Dun from the South Fork Sauk River in Washington
The lack of a darkened humeral crossvein rules out Epeorus albertae and Epeorus dulciana. The lack of a dark macula on the forefemora rules out Epeorus longimanus. The small size rules out Epeorus grandis and Epeorus permagnus. That leaves as the only possibility known in Washington state Epeorus deceptivus. It is a small species, although not reportedly quite as small as this specimen. I couldn't find anything in the species description in Traver (1935) to definitively confirm or rule out the species ID, given that I don't have the preserved specimen to check under a microscope, but it has to be either deceptivus or something not yet reported in Washington.

It was collected at the same time as a similar-sized female dun.
Epeorus vitreus (Heptageniidae) (Sulphur) Mayfly Nymph from unknown in Wisconsin

Mayfly Species Rhithrogena futilis

These are sometimes called Western Gordon Quills.
This uncommon and slightly larger species hatches earlier than Rhithrogena hageni, but otherwise is virtually indistinguishable.

Mayfly Species Epeorus albertae

These are very rarely called Western Gordon Quills.
This is one of the two most common Western species of Epeorus, and its spinner falls can be important.
Dorsal view of a Epeorus albertae (Heptageniidae) (Pink Lady) Mayfly Nymph from the East Fork Issaquah Creek in Washington
This specimen keys to the Epeorus albertae group of species. Of the five species in that group, the two known in Washington state are Epeorus albertae and Epeorus dulciana. Of the two, albertae has been collected in vastly more locations in Washington than dulciana, suggesting it is far more common. On that basis alone I'm tentatively putting this nymph in albertae, with the large caveat that there's no real information to rule out dulciana.

Mayfly Species Epeorus longimanus

These are very rarely called Western Gordon Quills.
Epeorus longimanus and Epeorus albertae are the two most common Epeorus species in the West.
Lateral view of a Male Epeorus longimanus (Heptageniidae) (Slate Brown Dun) Mayfly Dun from Mystery Creek #295 in Washington
Identification of this one was as follows. Body 9 mm, wing 11 mm.
Both Epeorus albertae and Epeorus dulciana should have a conspicuously darkened humeral crossvein in the forewing. This one doesn't.
The foretarsal claws are dissimilar (one sharp, one blunt), which also rules out the Epeorus albertae group.
The dark macula on the forefemora rules out Epeorus deceptivus, which is also supposed to be a little bit smaller.
Both Epeorus grandis and Epeorus permagnus should be much, much larger.
Of the species known to be present in Washington, this leaves only Epeorus longimanus, which is exactly the right size. The key to male spinners in Traver (1935) describes distinctive markings that are visible (although more faintly) in this dun: “Black posterior margins of tergites do not extend laterally to pleural fold, but an oblique black line form this margin cuts across poster-lateral triangle to pleural fold.”
Female Epeorus longimanus (Heptageniidae) (Slate Brown Dun) Mayfly Spinner from the Touchet River in Washington
Epeorus longimanus (Heptageniidae) (Slate Brown Dun) Mayfly Nymph from the Vermillion River in Montana

Western Gordon Quills

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