Jere -
Please straighten me out. I have never seen a big PMD. Early in the Summer they might be a #14, but more than likely a #16. and the next generations are quite small, like now...an #18 at the biggest. Maybe we are talking about a different bug.
PMD as a name is perhaps second only to Blue -winged Olive when it comes to a common name causing confusion for western anglers. Unfortunately, while some of the confusion can be resolved, some can't. What can be reasonably assumed accurate at this point is the following:
1. The common name "Pale Morning Dun" originally referred to quite a few pale western ephemerellids that have since been combined, "distilled" if you will, down to two species -
E. dorothea infrequens (prev.
E. infrequens and others) and
E. excrucians (prev.
E. inermis and others)
2. The two are virtually impossible for anglers to tell apart taxonomically. Neither size or certain nymphal characters are as dependable as once thought.
3. They do not have "broods" as do the baetids. They coexist as disparate species and populations with some strains smaller and later than others.
4. Both species can be as large as size 12 and at least
excrucians can be as small as size 18, though size 14 and 16 are the most common. I have fished over size 12 PMD's many times in the Sierra's (particularly one lake with a nice feeder coming into it where they come off in early June)
What can't be resolved at this time but assumed to be probable (in other words, the jury is still out):
1. Where both are found together, the largest and earliest of the two for a given watershed is
E. d. infrequens. It is also thought to be the shortest in duration.
2. The smaller, later, and longer lasting of the two is E. excrucians.
3. Alleged sightings of "PMD's" size 20 and smaller are probably false. The sizes may be exaggerated, anglers may be referring to the smaller sized flies that are often more effective, or they could be confusing them with the tiny sulfur baetids
Centroptilum and
Procloeon that hatch in the Summer from similar habitat.
The "flies in the ointment" for the above assumptions are populations of large
excrucians have been documented as well as possibly populations of small
d. infrequens. There are populations of "PMD's" hard to place with either species (over a variety of characters). Also, while not currently reported to be so tiny, there is documentation of other ephemerellids being as small as size 20, so who knows? Bottom line is there's a lot left to learn about these pale western ephemerellids.
Hope this helps.
Westco - What's not clear to me is whether you were at the river or not at the times that Dubbin reported. Besides having to be there more than 30 days ago, you had to be there pretty early as well if I read him right. I've never heard of them coming off at the crack of dawn before, usually running into them either during the day or right at dark on the blue-bird hot days at the end of their cycle. But, perhaps that's why I've missed them so many times.:) Regardless, I agree with Sayfu that Drakes are spotty as hell.