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Artistic view of a Male Pteronarcys californica (Pteronarcyidae) (Giant Salmonfly) Stonefly Adult from the Gallatin River in Montana
Salmonflies
Pteronarcys californica

The giant Salmonflies of the Western mountains are legendary for their proclivity to elicit consistent dry-fly action and ferocious strikes.

Dorsal view of a Setvena wahkeena (Perlodidae) (Wahkeena Springfly) Stonefly Nymph from Mystery Creek #199 in Washington
As far as I can tell, this species has only previously been reported from one site in Oregon along the Columbia gorge. However, the key characteristics are fairly unmistakable in all except for one minor detail:
— 4 small yellow spots on frons visible in photos
— Narrow occipital spinule row curves forward (but doesn’t quite meet on stem of ecdysial suture, as it's supposed to in this species)
— Short spinules on anterior margin of front legs
— Short rposterior row of blunt spinules on abdominal tergae, rather than elongated spinules dorsally
I caught several of these mature nymphs in the fishless, tiny headwaters of a creek high in the Wenatchee Mountains.
27" brown trout, my largest ever. It was the sub-dominant fish in its pool. After this, I hooked the bigger one, but I couldn't land it.
Troutnut is a project started in 2003 by salmonid ecologist Jason "Troutnut" Neuswanger to help anglers and fly tyers unabashedly embrace the entomological side of the sport. Learn more about Troutnut or support the project for an enhanced experience here.

Identification: Key to Species of Agnetina Adults

Identification: Key to Species of Agnetina Adults

Adapted from Stark et al (1986)
Option 1Option 2
Femora transversely banded
Anterior view of femur of Agnetina annulipes.
Anterior view of femur of Agnetina annulipes. Figure source: Stark et al (1986)
Femora longitudinally banded
Dorsal view of middle and hind femora of an Agnetina capitata male.
Dorsal view of middle and hind femora of an Agnetina capitata male. Figure from this Male Agnetina capitata adult.
Anterior view of femur of Agnetina capitata.
Anterior view of femur of Agnetina capitata. Figure source: Stark et al (1986)
Anterior view of femur of Agnetina flavescens.
Anterior view of femur of Agnetina flavescens. Figure source: Stark et al (1986)
Male hemitergal process foot-shaped with long slender ankle and toe
Lateral view of hemitergal process of male Agnetina annulipes.
Lateral view of hemitergal process of male Agnetina annulipes. Figure source: Stark et al (1986)
Male hemitergal process either with toe blunt or ankle short
Lateral view of hemitergal process of male Agnetina capitata.
Lateral view of hemitergal process of male Agnetina capitata. Figure source: Stark et al (1986)
Lateral view of hemitergal process of male Agnetina flavescens.
Lateral view of hemitergal process of male Agnetina flavescens. Figure source: Stark et al (1986)
Female subgenital plate truncate, dark pigment covers sternum 8 in continuous mesal band
Sterna 8 and 9 of female Agnetina annulipes.
Sterna 8 and 9 of female Agnetina annulipes. Figure source: Stark et al (1986)
Female subgenital plate rounded with areas of basolateral dark pigment
Sterna 8 and 9 of female Agnetina capitata.
Sterna 8 and 9 of female Agnetina capitata. Figure source: Stark et al (1986)
Sterna 8 and 9 of female Agnetina flavescens.
Sterna 8 and 9 of female Agnetina flavescens. Figure source: Stark et al (1986)
Remaining species: Agnetina capitata and Agnetina flavescens
2 Example Specimens
Agnetina annulipes Go to Couplet 2
Adapted from Stark et al (1986)
The current couplet is highlighted with darker colors and a icon, and couplets leading to this point have a icon.
Couplet 1

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Couplet 1 (You are here)
Leads to Agnetina annulipes:
  • Femora transversely banded
  • Male hemitergal process foot-shaped with long slender ankle and toe
  • Female subgenital plate truncate, dark pigment covers sternum 8 in continuous mesal band
Leads to Couplet 2:
  • Femora longitudinally banded
  • Male hemitergal process either with toe blunt or ankle short
  • Female subgenital plate rounded with areas of basolateral dark pigment
Couplet 2
Leads to Agnetina capitata:
  • Ventral margin of femora dark forming mesal yellow window
  • Apical portion of male hemitergal process long and sinuate, toe bluntly rounded
  • Dark pirgment areas on female sterna 8 and 9 large quadrangular
Leads to Agnetina flavescens:
  • Ventral margin of femora light
  • Swollen area of male hemitergal process adjacent to foot obscuring ankle area
  • Dark pigment areas of female sterna 8 and 9 not quadrangular

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References

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