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Lateral view of a Female Hexagenia limbata (Ephemeridae) (Hex) Mayfly Dun from the Namekagon River in Wisconsin
Hex Mayflies
Hexagenia limbata

The famous nocturnal Hex hatch of the Midwest (and a few other lucky locations) stirs to the surface mythically large brown trout that only touch streamers for the rest of the year.

Dorsal view of a Setvena wahkeena (Perlodidae) (Wahkeena Springfly) Stonefly Nymph from Mystery Creek #199 in Washington
As far as I can tell, this species has only previously been reported from one site in Oregon along the Columbia gorge. However, the key characteristics are fairly unmistakable in all except for one minor detail:
— 4 small yellow spots on frons visible in photos
— Narrow occipital spinule row curves forward (but doesn’t quite meet on stem of ecdysial suture, as it's supposed to in this species)
— Short spinules on anterior margin of front legs
— Short rposterior row of blunt spinules on abdominal tergae, rather than elongated spinules dorsally
I caught several of these mature nymphs in the fishless, tiny headwaters of a creek high in the Wenatchee Mountains.
27" brown trout, my largest ever. It was the sub-dominant fish in its pool. After this, I hooked the bigger one, but I couldn't land it.
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Mayfly Species Iswaeon anoka (Tiny Blue-Winged Olives)

This tiny hind-wingless Midwestern and Western species can produce excellent hatches because it is so extremely abundant. Its bright green duns are unmistakable. In the West they can also be an equally unmistakable bright almost fluorescent chartreuse, especially as nymphs. They are very common in the Northwest and Rocky Mountain states, with their population densities giving way to the similarly tiny and hind-wingless (though more somber colored) Acentrella species in Southern Oregon, California and the Southwest.

Iswaeon anoka was first brought to the attention of the angling community (as Pseudocloeon anoka) by famous angling author and columnist Joe Brooks. Back in the late 60's he wrote an article for Outdoor Life magazine extolling the work of Doug Swisher and Carl Richards in their forthcoming and groundbreaking book, Selective Trout. This species was featured as the model for their now famous version of the "No-Hackle" dry fly and imitative parachute patterns, sparking a revolution in fly design for hyper-selective trout. The rest as they say, is history...

Taxonomic History

This species appears in some angling literature under the former names Pseudocloeon anoka in the Midwest and Pseudocloeon edmundsi in the West.

Where & when

Time of year : Late June through late July; again from August through October

In 10 records from GBIF, adults of this species have been collected during July (40%), May (30%), June (20%), and September (10%).

In 4 records from GBIF, this species has been collected at elevations of 3, 26, 886, and 1089 ft.

Species Range

Hatching behavior

Fred Arbona says of Iswaeon anoka (as Pseudocloeon anoka) hatching behavior in Mayflies, the Angler, and the Trout that:

The little nymph can easily crawl out of the water during emergence, and therefore the trout are able to consume them en masse very close to the bank.


This is somewhat open to interpretation and it is suspected they emerge on the surface as well.

Nymph biology

Substrate: Gravel, vegetation

The nymphs are reported to prefer shallow water near the banks.

Identification

Resources

There's a great blog post on Aquatic Insects of Central Virginia about identifying Iswaeon anoka.


Start a Discussion of Iswaeon anoka

References

Mayfly Species Iswaeon anoka (Tiny Blue-Winged Olives)

Taxonomy
Species Range
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