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Artistic view of a Male Pteronarcys californica (Pteronarcyidae) (Giant Salmonfly) Stonefly Adult from the Gallatin River in Montana
Salmonflies
Pteronarcys californica

The giant Salmonflies of the Western mountains are legendary for their proclivity to elicit consistent dry-fly action and ferocious strikes.

Dorsal view of a Setvena wahkeena (Perlodidae) (Wahkeena Springfly) Stonefly Nymph from Mystery Creek #199 in Washington
As far as I can tell, this species has only previously been reported from one site in Oregon along the Columbia gorge. However, the key characteristics are fairly unmistakable in all except for one minor detail:
— 4 small yellow spots on frons visible in photos
— Narrow occipital spinule row curves forward (but doesn’t quite meet on stem of ecdysial suture, as it's supposed to in this species)
— Short spinules on anterior margin of front legs
— Short rposterior row of blunt spinules on abdominal tergae, rather than elongated spinules dorsally
I caught several of these mature nymphs in the fishless, tiny headwaters of a creek high in the Wenatchee Mountains.
27" brown trout, my largest ever. It was the sub-dominant fish in its pool. After this, I hooked the bigger one, but I couldn't land it.
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Caddisfly Species Brachycentrus americanus (American Grannoms)

Where & when

Time of year : July and August

This species is most prolific in the West and Midwest, but it may still be of concern to fishermen in the East.

In 80 records from GBIF, adults of this species have mostly been collected during July (31%), August (28%), June (16%), September (13%), and May (9%).

In 109 records from GBIF, this species has been collected at elevations ranging from 3 to 10007 ft, with an average (median) of 6099 ft.

Species Range

Hatching behavior

Time of day : Morning

Special thanks to Lloyd Gonzales, author of the excellent new book Fly-Fishing Pressured Water, for helping to sort out the Brachycentrus species and common names.

Specimens of the Caddisfly Species Brachycentrus americanus

1 Female Adult
2 Larvae

Discussions of Brachycentrus americanus

Brachycentrus americanus on the Lower Sacramento River California
Posted by Troutguide on Oct 29, 2016
Last reply on Oct 29, 2016 by Troutguide
I believe this is the species found in sometimes very large numbers on the Lower Sacramento River in the Redding area. Ten years ago it was present in such large numbers that fishing a fly on the bottom resulted in frequently hooking one of these caddis still in its case. Along with other aquatic insects their numbers have declined to a fraction of once seen. I don't believe the egg Sac dropped by the females to be olive , instread I have seen it to be a bright green. The females seem to oviposit close to the edge of flowing water and not midstream.

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References

Caddisfly Species Brachycentrus americanus (American Grannoms)

Taxonomy
Species Range
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