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Lateral view of a Female Hexagenia limbata (Ephemeridae) (Hex) Mayfly Dun from the Namekagon River in Wisconsin
Hex Mayflies
Hexagenia limbata

The famous nocturnal Hex hatch of the Midwest (and a few other lucky locations) stirs to the surface mythically large brown trout that only touch streamers for the rest of the year.

Dorsal view of a Grammotaulius betteni (Limnephilidae) (Northern Caddisfly) Caddisfly Larva from the Yakima River in Washington
This is a striking caddis larva with an interesting color pattern on the head. Here are some characteristics I was able to see under the microscope, but could not easily expose for a picture:
- The prosternal horn is present.
- The mandible is clearly toothed, not formed into a uniform scraper blade.
- The seems to be only 2 major setae on the ventral edge of the hind femur.
- Chloride epithelia seem to be absent from the dorsal side of any abdominal segments.
Based on these characteristics and the ones more easily visible from the pictures, this seems to be Grammotaulius. The key's description of the case is spot-on: "Case cylindrical, made of longitudinally arranged sedge or similar leaves," as is the description of the markings on the head, "Dorsum of head light brownish yellow with numerous discrete, small, dark spots." The spot pattern on the head is a very good match to figure 19.312 of Merritt R.W., Cummins, K.W., and Berg, M.B. (2019). The species ID is based on Grammotaulius betteni being the only species of this genus known in Washington state.
27" brown trout, my largest ever. It was the sub-dominant fish in its pool. After this, I hooked the bigger one, but I couldn't land it.
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Stonefly Species Isoperla miwok (Yellow Sallies)

Species Range

Identification

Source: The Isoperla Of California (Plecoptera: Perlodidae); Larval Descriptions And A Key To 17 Western Nearctic Species

Diagnosis. Male larvae of Isoperla miwok are distinctive among all other California Isoperla species with 2 – 4 total submarginal (A + B) setae by the single lacinia submarginal (A) seta arising from near the middle of the base of the subapical tooth (Fig. 11 h).

Physical description

Most physical descriptions on Troutnut are direct or slightly edited quotes from the original scientific sources describing or updating the species, although there may be errors in copying them to this website. Such descriptions aren't always definitive, because species often turn out to be more variable than the original describers observed. In some cases, only a single specimen was described! However, they are useful starting points.

Source: The Isoperla Of California (Plecoptera: Perlodidae); Larval Descriptions And A Key To 17 Western Nearctic Species

Male larva. Body length of mature larva 9 – 11 mm. Dorsum of head with contrasting pigment pattern and fine light clothing setae, anterior frontoclypeus margin unpigmented; light M shaped pattern anterior to median ocellus variable, usually connected to light frontoclypeus area by an apically narrowed median longitudinal light band, lateral thin arms directed posterolaterally, extending to antennal bases; posterior ocelli with partially enclosed large light areas along outer lateral margins; interocellar area variable, usually partially light, completely enclosed by dark pigment and extending posteriorly to epicranial Y-suture or sometimes open posteriorly and connected to posterior margin of head by thin light longitudinal band; occiput with irregular spinulae band extending from below eye to near median epicranial suture, not enclosed completely by dark pigment (Fig. 11 a). Lacinia bidentate, total length 622 – 888 µm (Figs. 2 i, 11 e- h, Table 2 - 4); submarginal row (A + B) with 2 – 3 setae, groups A-B interrupted by gap below subapical tooth (SAT) inner margin (Fig. 11 g); 1 submarginal seta (A) inserted near mid-base of SAT, half as stout at its base as first submarginal (B) seta, plus 1 thin marginal seta (TMS) adjacent to apical tooth (AT) inner margin, sometimes obstructed from view by AT or broken, and 1 dorsal seta (DS) located below SAT inner margin, partially obstructed by SAT (Figs. 11 g-h); 1 – 2 submarginal setae (B) located past SAT inner margin (Fig. 11 h); 5 – 7 marginal setae (C), initially long-stout and widely spaced, last few shorter and closer, blending into and difficult to differentiate from dorsal surface setae (Fig. 11 e); 7 – 15 ventral surface setae (D), scattered below marginal setae, ending posteriorly at approximately ¾ the inner lacinia margin length and concentrated in posterior half (Fig. 11 f); dorsal surface setae (DSS) continue from last marginal setae (C) as a single, laterally protruding, submarginal row (sometimes 2 – 3 setae thick) along inner-lateral margin, ending before or near posterior-most ventral surface setae (Fig. 11 f); 10 – 15 minute scale-like spinulae rarely present on and along ventral inner-lateral margin at middle of lacinia. Galea with 12 – 26 setae in sparse ventral row, apex with 2 – 4 setae. Maxillary Palp segments 2 – 3 with curved, apically pointed setae. Pronotum with large median light area bordered by thick dark comma shaped bands (typical of the Isoperla sobria complex) that fade rapidly in alcohol; discs each with fine light clothing setae, except over small light rugosites and lateral margins with broad light bands (Fig. 11 b); Mesonotum and metanotum with contrasting pigment pattern and fine light clothing setae (Fig. 11 c). Legs with numerous fine light clothing setae and scattered erect spines on outer surface of femora, erect spines longest and concentrated along dorsal surface; fine silky setae numerous and continuous on dorsal surfaces of femora and tibia (Fig. 20 i); distal femora and proximal tibia with faint transverse bands. Abdominal terga with three distinct longitudinal dark stripes; wide median longitudinal light band bisected with thin dark median longitudinal stripe; lateral pair of dark longitudinal stripes about as wide as median dark stripe, not extending to lateral margins; numerous fine light clothing setae and erect spines scattered dorsally; posterior margin with scattered long and numerous short spines in a concentrated row (Fig. 11 d).

Source: The Isoperla Of California (Plecoptera: Perlodidae); Updated Male Descriptions And Adult Keys For 18 Western Nearctic Species

Male. Aedeagus: sclerotized posterior process absent; body with one small posterior lobe, one large dorsal lobe and a pair of small pointed lobes on apicolateral margins (Fig. 9 a-b); a pair of long curved patches of spinulae on posteroventral surface (Fig. 9 b), and a broad patch of spinules on the posteroapical surface (Figs. 9 a-c). Abdominal terga 8 - 9, 9, 9 - 10: without stout spinulae or long stout setae. Posterolateral margins of at least abdominal segment 8 with scale-like setae clustered in brushes of several setae. Paraprocts: curved dorsally, length if straightened subequal to combined first and second cercal segments, tapering abruptly to blunt apices (Fig. 9 c). Vesicle: lobe barely perceptible, vestigial when present, widest at base with broadly rounded apex (Fig. 9 d).


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References

Stonefly Species Isoperla miwok (Yellow Sallies)

Species Range
Resources
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