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Lateral view of a Female Hexagenia limbata (Ephemeridae) (Hex) Mayfly Dun from the Namekagon River in Wisconsin
Hex Mayflies
Hexagenia limbata

The famous nocturnal Hex hatch of the Midwest (and a few other lucky locations) stirs to the surface mythically large brown trout that only touch streamers for the rest of the year.

Dorsal view of a Setvena wahkeena (Perlodidae) (Wahkeena Springfly) Stonefly Nymph from Mystery Creek #199 in Washington
As far as I can tell, this species has only previously been reported from one site in Oregon along the Columbia gorge. However, the key characteristics are fairly unmistakable in all except for one minor detail:
— 4 small yellow spots on frons visible in photos
— Narrow occipital spinule row curves forward (but doesn’t quite meet on stem of ecdysial suture, as it's supposed to in this species)
— Short spinules on anterior margin of front legs
— Short rposterior row of blunt spinules on abdominal tergae, rather than elongated spinules dorsally
I caught several of these mature nymphs in the fishless, tiny headwaters of a creek high in the Wenatchee Mountains.
27" brown trout, my largest ever. It was the sub-dominant fish in its pool. After this, I hooked the bigger one, but I couldn't land it.
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Stonefly Species Isoperla marmorata (Yellow Sallies)

Species Range

Identification

Source: The Isoperla Of California (Plecoptera: Perlodidae); Larval Descriptions And A Key To 17 Western Nearctic Species

Diagnosis. Male larvae of the Isoperla marmorata complex were differentiated from other species with 1 – 2 lacinia submarginal (A) setae by having 41 – 110 ventral surface setae and pronotal discs mostly dark. Within the complex, Isoperla marmorata was distinguished from Isoperla roguensis by usually having the light M shaped pattern connected to the light anterior frontoclypeus area by a light longitudinal band (See also diagnosis under Isoperla fulva). The only exception to the light M shaped pattern character occurred in 8 exuviae collected from Prairie Creek, Humboldt County, California. These possessed partial to entire M shaped light patterns, but were not connected to the light anterior frontal clypeus area.

Physical description

Most physical descriptions on Troutnut are direct or slightly edited quotes from the original scientific sources describing or updating the species, although there may be errors in copying them to this website. Such descriptions aren't always definitive, because species often turn out to be more variable than the original describers observed. In some cases, only a single specimen was described! However, they are useful starting points.

Source: The Isoperla Of California (Plecoptera: Perlodidae); Larval Descriptions And A Key To 17 Western Nearctic Species

Male larva. Body length of mature larva 11 – 13 mm. Dorsum of head with contrasting pigment pattern and fine dark clothing setae, anterior frontoclypeus margin unpigmented; light M shaped pattern anterior to median ocellus indistinct, connected (sometimes partially) to light frontoclypeus area by a wide median longitudinal light band, lateral thin arms directed posterolaterally, extending to antennal bases; posterior ocelli with partially enclosed large light areas along outer lateral margins; interocellar area variable, usually partially light, completely enclosed by dark pigment and not extending past posterior ocelli; occiput with irregular spinulae band extending from below eye to near median epicranial suture, not enclosed completely by dark pigment (Fig. 10 a). Lacinia bidentate, total length 816 – 974 µm (Figs. 2 h, 10 a-h, Tables 2 - 4); submarginal row (A + B) with 2 – 3 setae, groups A-B interrupted by gap below subapical tooth (SAT) inner margin (Fig. 10 g); 1 submarginal seta (A) inserted at base of apical tooth (AT) inner margin, plus 1 thin marginal seta (TMS) adjacent to AT inner margin, sometimes obstructed from view by AT, submarginal seta (A) or broken, and 1 dorsal seta (DS) located below SAT inner margin, partially obstructed by SAT or submarginal setae (B) (Figs. 10 g-h); 1 – 2 submarginal setae (B) located past SAT inner margin (Fig. 10 h); 7 – 10 marginal setae (C), initially long-stout and widely spaced, last few shorter and variably spaced, blending into and difficult to differentiate from dorsal and ventral surface setae (Fig. 10 e); 72 – 110 ventral surface setae (D) forming dense longitudinal band below submarginal and marginal setae, ending posteriorly at approximately ¾ the inner lacinia margin length, setae closest to inner margin protrude laterally past lacinia margin (Fig. 10 f); dorsal surface setae (DSS) forming dense, laterally protruding, longitudinal band on and along inner-lateral margin, ending before posterior-most ventral surface setae (Fig. 10 f). Galea with 19 – 40 setae in sparse ventral row, apex with 4 – 5 setae. Maxillary Palp segments 2 – 3 with curved, apically pointed setae. Pronotum with median light area bordered by wide dark bands typical of the Isoperla marmorata complex; discs each with light rugosites concentrated along median dark margins and fine dark clothing setae, lateral margins with broad light bands (Fig. 10 b). Mesonotum and metanotum with contrasting pigment pattern and fine dark clothing setae (Fig. 10 c). Legs with numerous fine golden clothing setae and scattered erect spines on outer surface of femora, erect spines longest and concentrated on dorsal surface; fine silky setae sparse on dorsal surface of femora, numerous and continuous on tibia (Fig. 20 h); tibia with faint transverse bands near proximal end. Abdominal terga variable, usually with two distinct longitudinal dark stripes; wide light median longitudinal band sometimes bisected with faint, light brown longitudinal stripe; lateral pair of dark longitudinal stripes usually not extending to lateral margins; numerous fine dark clothing setae and erect spines scattered dorsally; posterior margin with scattered long and numerous short spines in a concentrated row (Fig. 10 d).

Source: The Isoperla Of California (Plecoptera: Perlodidae); Updated Male Descriptions And Adult Keys For 18 Western Nearctic Species

Male. Aedeagus: sclerotized posterior process present; body with one small posterior lobe, one dorsal lobe and an anterior lobe (Fig. 8 a); one small patch of spinulae concentrated on the inverted membrane above sclerotized process (Figs. 8 a-b); sclerotized process approaching 0.5 mm when fully extended, blade-like in lateral view (Fig. 8 a), distal margin thin in posterior view (Fig. 8 b), and narrow proximal portion of sclerotized process usually inverted inside posterior aedeagal membrane. Abdominal terga 8 - 9, 9, 9 - 10: without stout spinulae or long stout setae. Posterolateral margins of at least abdominal segment 8 with scale-like setae clustered in brushes of several setae. Paraprocts: curved dorsally, length if straightened subequal to combined first and second cercal segments, tapering gradually to long blunt apices (Fig. 8 c). Vesicle: rounded lobe, widest at base with broadly rounded apical margin (Fig. 8 d).


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Stonefly Species Isoperla marmorata (Yellow Sallies)

Species Range
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