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Artistic view of a Male Pteronarcys californica (Pteronarcyidae) (Giant Salmonfly) Stonefly Adult from the Gallatin River in Montana
Salmonflies
Pteronarcys californica

The giant Salmonflies of the Western mountains are legendary for their proclivity to elicit consistent dry-fly action and ferocious strikes.

Dorsal view of a Holocentropus (Polycentropodidae) Caddisfly Larva from the Yakima River in Washington
This one seems to tentatively key to Holocentropus, although I can't make out the anal spines in Couplet 7 of the Key to Genera of Polycentropodidae Larvae nor the dark bands in Couplet 4 of the Key to Genera of Polycentropodidae Larvae, making me wonder if I went wrong somewhere in keying it out. I don't see where that could have happened, though. It might also be that it's a very immature larva and doesn't possess all the identifying characteristics in the key yet. If Holocentropus is correct, then Holocentropus flavus and Holocentropus interruptus are the two likely possibilities based on range, but I was not able to find a description of their larvae.
27" brown trout, my largest ever. It was the sub-dominant fish in its pool. After this, I hooked the bigger one, but I couldn't land it.
Troutnut is a project started in 2003 by salmonid ecologist Jason "Troutnut" Neuswanger to help anglers and fly tyers unabashedly embrace the entomological side of the sport. Learn more about Troutnut or support the project for an enhanced experience here.

Olive Quills

Like most common names,"Olive Quill" can refer to more than one taxon. They're previewed below, along with 3 specimens. For more detail click through to the scientific names.

Mayfly Species Ephemerella excrucians

These are sometimes called Olive Quills.
For trout (if not anglers), this single species is arguably the most important mayfly in North America. In terms of sheer numbers, breadth of distribution and hatch duration, it has a good argument.

Ephemerella excrucians or Pale Morning Dun usually follows its larger sibling Ephemerella dorothea infrequens with which it shares the same common name. What it often lacks in size by comparison is made up for with it's duration, often lasting for months with intermittent peaks. This close relationship with infrequens has led many anglers to confuse Pale Morning Dun biology with that of the multivoltine Baetidae species, having disparate broods that decrease in size as the season advances. Sharing the same common name has not helped to alleviate this misconception.

Until recently, Ephemerella excrucians was considered primarily an upper MidWestern species of some regional importance commonly called Little Red Quill among other names. Recent work by entomologists determined that it is actually the same species as the important Western Pale Morning Dun (prev.Ephemerella inermis), and the lake dwelling Sulphur Dun of the Yellowstone area, (prev.Ephemerella lacustris). Since all three are considered variations of the same species, they have been combined into excrucians, being the original name for the type species reported as far back as the Civil War. Angler speculation had simmered for some time that the stillwater loving Ephemerella lacustris was much more widespread, inhabiting more water types then previously thought and could account for many large sulfurish ephemerellids found in still to very slow water locations throughout the West. With the revisions, this discussion is now moot.

Ephemerella excrucians variability in appearance, habitat preferences, and wide geographical distribution are cause for angler confusion with the changes in classification. They can be pale yellow 18's on a large Oregon river, creamy orange 14's on western lakes and feeder streams, large olive green on CA spring creeks as well as tiny sulfur ones in many Western watersheds. Then there's the little Red Quill on small streams in Wisconsin. Yet, all are the same species.
Lateral view of a Female Ephemerella excrucians (Ephemerellidae) (Pale Morning Dun) Mayfly Spinner from the Bois Brule River in Wisconsin
Dorsal view of a Ephemerella excrucians (Ephemerellidae) (Pale Morning Dun) Mayfly Nymph from the Bois Brule River in Wisconsin
I spent a while with a microscope to fairly positively identify this specimen as Ephemerella excrucians.

Mayfly Species Serratella serrata

These are sometimes called Olive Quills.
This locally important species is rarely mentioned in fly fishing literature, and what little information is given is identical to that for Teloganopsis deficiens. Knopp and Cormier say both species can produce good hatches.

Mayfly Species Drunella walkeri

These are very rarely called Olive Quills.
Ernest Schwiebert had this to say about the hatches of this chunky Eastern Drunella (Blue-Winged Olive) species:

An imitation is rarely required, but notes show that when it is needed it is needed badly.


See the Drunella and Drunella cornuta hatch pages for additional information.

References

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