This one was very puzzling to identify. It's clearly Heptageniidae
. It keys pretty easily to couplet 58 in Merritt, Cummins, & Berg 5th Edition, but that's where things get tricky.
58: Male penes
distinctly L-shaped; Segment 1 of foretarsi
usually 1/3 to 2/3 the length of segment 2
58': Males penes
not distinctly L-shaped; Segment 1 of foretarsi
usually 1/5 to 1/2 the length of segment 2
The characteristics conflict here, as I'd say the penes
ARE L-shaped, but the fore tarsal
ratio is less than 1/3. Maybe the weasel word "usually" allows for that, and it's some kind of Stenonema
, for which I'm not aware of any records in Montana, let alone western Montana.
Going by the tarsal
ratio and assuming it's not Stenonema
, everything in couplet 60 points to Afghanurus
: contiguous compound eyes
, genitalia similar to fig 13.222 in MC&B, and weakly developed basal costal crossveins
in the forewings. But I looked up the original descriptions of the three Afghanurus
species, and the body sizes (flowersi 5-7 mm, inconspicua 4 mm, joernensis 6-7 mm) are way below this one's 10 mm.
This led me to notice that the closely related Ecdyonurus
is not included in the key at all, so I checked the descriptions of those species and found that Ecdyonurus criddlei
fits very well: the color of the legs, the markings on the tergites
, the short fore tarsus
and ratios of the segments, etc. It was originally described as 7 mm, smaller than the 10 mm body of this specimen, but synonyms
have been described with larger sizes including wing lengths up to 11 mm for females for Heptagenia salvini
This specimen was found with a female
nearby, which I think is probably the same species, although it is difficult to be certain.