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Lateral view of a Female Hexagenia limbata (Ephemeridae) (Hex) Mayfly Dun from the Namekagon River in Wisconsin
Hex Mayflies
Hexagenia limbata

The famous nocturnal Hex hatch of the Midwest (and a few other lucky locations) stirs to the surface mythically large brown trout that only touch streamers for the rest of the year.

Dorsal view of a Grammotaulius betteni (Limnephilidae) (Northern Caddisfly) Caddisfly Larva from the Yakima River in Washington
This is a striking caddis larva with an interesting color pattern on the head. Here are some characteristics I was able to see under the microscope, but could not easily expose for a picture:
- The prosternal horn is present.
- The mandible is clearly toothed, not formed into a uniform scraper blade.
- The seems to be only 2 major setae on the ventral edge of the hind femur.
- Chloride epithelia seem to be absent from the dorsal side of any abdominal segments.
Based on these characteristics and the ones more easily visible from the pictures, this seems to be Grammotaulius. The key's description of the case is spot-on: "Case cylindrical, made of longitudinally arranged sedge or similar leaves," as is the description of the markings on the head, "Dorsum of head light brownish yellow with numerous discrete, small, dark spots." The spot pattern on the head is a very good match to figure 19.312 of Merritt R.W., Cummins, K.W., and Berg, M.B. (2019). The species ID is based on Grammotaulius betteni being the only species of this genus known in Washington state.
27" brown trout, my largest ever. It was the sub-dominant fish in its pool. After this, I hooked the bigger one, but I couldn't land it.
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Mayfly Species Callibaetis fluctuans (Speckled Duns)

This uncommon species may be locally abundant.

Where & when

In 24 records from GBIF, adults of this species have mostly been collected during August (38%), July (29%), September (17%), and June (13%).

In 20 records from GBIF, this species has been collected at elevations ranging from 325 to 7306 ft, with an average (median) of 3337 ft.

Species Range

Physical description

Most physical descriptions on Troutnut are direct or slightly edited quotes from the original scientific sources describing or updating the species, although there may be errors in copying them to this website. Such descriptions aren't always definitive, because species often turn out to be more variable than the original describers observed. In some cases, only a single specimen was described! However, they are useful starting points.

Female Spinner

Body length: 6-7 mm
Wing length: 6.5-8 mm

A brownish white species; cross veins of fore wing relatively few in number; marginal intercalaries single; wing of male not pigmented.

Head and thorax brownish white; vertex of female sometimes ferruginous, “with a double longitudinal carina.” Antennal filament dusky, sometimes pale at the base, a double light brown band on the thoracic notum, confluent behind. Legs whitish, the tips of the tarsi cloudy. Wings hyaline, iridescent; veins rather coarse near the base; longitudinal veins generally brown, and occasionally edged with brown toward the base. Cross veins fine, always hyaline. In the wing of the female, the space between costa and subcosta is hyaline; on the costal margin are 15 to 18 small brown spots, some of these confluent. A light brown band is present behind the subcosta, containing about 14 round hyaline spots—“some of them confluent before or behind with the hyaline part of the wing.” Posterior margin of this brown vitta variable, irregular; sometimes with 6 or 7 “large obtuse teeth.” Marginal intercalaries occur singly. Cross veins behind the vitta relatively few in number (14 to 18, according to Walsh).

Abdomen, both tergites and sternites, generally brownish-white, sometimes varied with brown. Two of Walsh’s specimens were pale brown, the 6th segment brown. Tails whitish. (Original description, of female only). The wing of the male (if we are correct in the identification of this species) is unpigmented. In both sexes, brown submedian curved marks are present on tergites and sternites.


Start a Discussion of Callibaetis fluctuans

References

Mayfly Species Callibaetis fluctuans (Speckled Duns)

Taxonomy
Species Range
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